List of crocidurines

Gray shrew
Greater white-toothed shrew (Crocidura russula)

Crocidurinae is a subfamily of small mammals in the shrew family Soricidae, which in turn is part of the order Eulipotyphla. A member of this family is called a crocidurine, or a white-toothed shrew. Crocidurinae is one of three subfamilies in Soricidae, along with the African shrews of Myosoricinae and the red-toothed shrews of Soricinae. They are found in Africa, Europe, and Asia, primarily in forests, savannas, shrublands, and grasslands, though some species can also be found in wetlands, rocky areas, or coastal areas. They range in size from the Etruscan shrew, one of the smallest mammal species at 3 cm (1 in) plus a 2 cm (1 in) tail,[1] to the hero shrew and Thor's hero shrew, at 15 cm (6 in) plus a 10 cm (4 in) tail. Crocidurines primarily eat invertebrates, small mammals, frogs, toads, and lizards. Almost no crocidurines have population estimates, but 21 species are categorized as endangered species, while the Andaman shrew, Christmas Island shrew, Harenna shrew, Jenkins's shrew, Nicobar shrew, and Wimmer's shrew are categorized as critically endangered.

The 235 extant species of Crocidurinae are divided into nine genera; 192 of them are in Crocidura, the most species of any mammal genus.[2] A few extinct prehistoric Crocidurinae species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[3]

Conventions

[edit]
IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically Endangered (6 species)
 EN Endangered (21 species)
 VU Vulnerable (12 species)
 NT Near threatened (7 species)
 LC Least concern (128 species)
Other categories
 DD Data deficient (60 species)
 NE Not evaluated (0 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the crocidurine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

[edit]

The subfamily Crocidurinae consists of nine genera: Crocidura, containing 192 species; Diplomesodon and Feroculus with a single species each; Paracrocidura, with three species; Ruwenzorisorex, with one species; Scutisorex, containing two species; Solisorex, with one species, Suncus, containing nineteen species; and Sylvisorex, containing fifteen species.

Subfamily Crocidurinae

  • Genus Crocidura (white-toothed shrews): 191 species
  • Genus Diplomesodon (piebald shrew): one species
  • Genus Feroculus (Kelaart's long-clawed shrew): one species
  • Genus Paracrocidura (large-headed shrews): three species
  • Genus Ruwenzorisorex (Ruwenzori shrew): one species
  • Genus Scutisorex (hero shrews): two species
  • Genus Solisorex (Pearson's long-clawed shrew): one species
  • Genus Suncus (musk shrews and pygmy shrews): nineteen species
  • Genus Sylvisorex (forest shrews): fifteen species
Crocidurinae[4]  

Crocidurines

[edit]

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[5]

Genus Crocidura Wagler, 1832 – 192 species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
African black shrew


C. nigrofusca
Matschie, 1895
Central and eastern Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[7]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[7]

African dusky shrew


C. caliginea
Hollister, 1916
Central Africa
Map of range
Size: 6–8 cm (2–3 in) long, plus 4–6 cm (2 in) tail[8]

Habitat: Forest[9]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[9]

African giant shrew

C. olivieri
(Lesson, 1827)

20 subspecies
  • C. o. anchietae
  • C. o. bueae
  • C. o. cara
  • C. o. cinereoaenea
  • C. o. darfurea
  • C. o. giffardi
  • C. o. guineensis
  • C. o. hansruppi
  • C. o. hedenborgiana
  • C. o. kivu
  • C. o. manni
  • C. o. martiensseni
  • C. o. nyansae
  • C. o. occidentalis
  • C. o. odorata
  • C. o. olivieri
  • C. o. spurelli
  • C. o. sururae
  • C. o. toritensis
  • C. o. zuleika
Sub-Saharan Africa
Map of range
Size: 11–14 cm (4–6 in) long, plus 8–10 cm (3–4 in) tail[10]

Habitat: Forest, savanna, and shrubland[11]

Diet: Invertebrates[10]
 LC 


Unknown Unknown[11]

Andaman shrew


C. andamanensis
Miller, 1902
South Andaman Island in India
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[12]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Population declining[12]

Andaman spiny shrew


C. hispida
Thomas, 1913
South Andaman Island in India
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[13]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown Unknown[13]

Annamite shrew


C. annamitensis
Jenkins, Lunde, & Moncrieff, 2009
Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[14]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[14]

Ansell's shrew


C. ansellorum
Hutterer & Dippenaar, 1987
South-central Africa
Map of range
Size: 5–6 cm (2–2 in) long, plus 4–6 cm (2 in) tail[15]

Habitat: Forest[16]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[16]

Arabian shrew


C. arabica
Hutterer, 1988
Oman and Yemen
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[17]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[17]

Armenian shrew


C. armenica
Gureev, 1963
Armenia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas[18]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[18]

Asian gray shrew


C. attenuata
H. Milne-Edwards, 1872
Eastern and southeastern Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[19]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[19]

Asian lesser white-toothed shrew

Brown shrew

C. shantungensis
Miller, 1901

Two subspecies
  • C. s. quelpartis
  • C. s. shantungensis
Eastern Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and grassland[20]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[20]

Bailey's shrew


C. baileyi
Osgood, 1936
Ethiopia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland[21]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population declining[21]

Bale shrew


C. bottegoides
Hutterer & Yalden, 1990
Ethiopia
Map of range
Size: 4–6 cm (2 in) long, plus 4–5 cm (2 in) tail[22]

Habitat: Grassland and forest[23]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[23]

Banka shrew


C. vosmaeri
Jentink, 1888
Bangka Island in Indonesia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[24]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[24]

Batak shrew


C. batakorum
Hutterer, 2007
Philippines
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[25]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[25]

Bates's shrew


C. batesi
(Dollman, 1915)
Western Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[26]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[26]

Beccari's shrew


C. beccarii
Dobson, 1886
Sumatra
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[27]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population declining[27]

Beletta shrew


C. yaldeni
Lavrenchenko, Voyta, & Hutterer, 2016
Ethiopia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[28]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[28]

Bicolored musk shrew


C. fuscomurina
(Heuglin, 1865)
Sub-Saharan Africa
Map of range
Size: 4–8 cm (2–3 in) long, plus 2–5 cm (1–2 in) tail[29]

Habitat: Savanna, grassland, and desert[30]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[30]

Bicolored shrew

Brown shrew

C. leucodon
(Hermann, 1780)
Europe and western Asia
Map of range
Size: 6–9 cm (2–4 in) long, plus 2–5 cm (1–2 in) tail[31]

Habitat: Shrubland and grassland[32]

Diet: Small mammals, frogs, toads, lizards, and invertebrates[31]
 LC 


Unknown Unknown[32]

Black-footed shrew

Gray shrew

C. nigripes
Miller & Hollister, 1921

Two subspecies
  • C. n. lipara
  • C. n. nigripes
Sulawesi island in Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[33]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[33]

Blackish white-toothed shrew


C. nigricans
Bocage, 1889
Angola
Map of range
Size: 6–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[34]

Habitat: Unknown[35]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[35]

Bornean shrew


C. foetida
Peters, 1870

Three subspecies
  • C. f. doriae
  • C. f. foetida
  • C. f. kelabit
Borneo
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[36]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[36]

Bottego's shrew


C. bottegi
Thomas, 1898
Ethiopia
Map of range
Size: 4–6 cm (2 in) long, plus 2–3 cm (1 in) tail[37]

Habitat: Unknown[38]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[38]

Buettikofer's shrew


C. buettikoferi
Jentink, 1888
West-central Africa
Map of range
Size: 7–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[39]

Habitat: Forest[40]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[40]

Butiaba naked-tailed shrew


C. littoralis
Heller, 1910
Central Africa
Map of range
Size: 8–10 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[41]

Habitat: Forest[42]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[42]

Cameroonian shrew


C. picea
Sanderson, 1940
West-central Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[43]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[43]

Canarian shrew

Gray shrew

C. canariensis
Hutterer, López-Jurado, & Vogel, 1987
Canary Islands
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Coastal marine, desert, and shrubland[44]

Diet: Snails and insects[44]
 EN 


Unknown Population declining[44]

Caspian shrew


C. caspica
Thomas, 1907
Iran and Azerbaijan
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[45]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[45]

Chinese white-toothed shrew


C. rapax
Allen, 1923

Four subspecies
  • C. r. kurodai
  • C. r. lutaoensis
  • C. r. rapax
  • C. r. tadae
Eastern Asia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[46]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[46]

Christmas Island shrew


C. trichura
Dobson, 1889
Christmas Island
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[47]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


0–200 Population declining[47]

Cinderella shrew


C. cinderella
Thomas, 1911
Western Africa
Map of range
Size: 5–7 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[48]

Habitat: Savanna[49]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[49]

Congo white-toothed shrew


C. congobelgica
Hollister, 1916
Central Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[50]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[50]

Cranbrook's white-toothed shrew


C. cranbrooki
Jenkins, Lunde, & Moncrieff, 2009
Northern Myanmar Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[51]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[51]

Cretan shrew


C. zimmermanni
Wettstein, 1953
Island of Crete in Greece
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown and rocky areas[52]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[52]

Crosse's shrew


C. crossei
Thomas, 1895
Western Africa
Map of range
Size: 6–7 cm (2–3 in) long, plus 4–6 cm (2 in) tail[53]

Habitat: Forest[54]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[54]

Cyrenaica shrew


C. aleksandrisi
Vesmanis, 1977
Northern Libya
Map of range
Size: 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[55]

Habitat: Shrubland[56]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[56]

Dent's shrew

Black and white picture of a schrew

C. denti
Dollman, 1915
Western and central Africa
Map of range
Size: About 6 cm (2 in) long, plus 5 cm (2 in) tail[57]

Habitat: Forest and savanna[58]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[58]

Desert musk shrew


C. smithii
Thomas, 1895

Two subspecies
  • C. s. debalsaci
  • C. s. smithii
Western and Eastern Africa
Map of range
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[59]

Habitat: Savanna[60]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[60]

Desperate shrew


C. desperata
Hutterer, Jenkins, & Verheyen, 1991
Tanzania
Map of range
Size: 8–11 cm (3–4 in) long, plus about 7 cm (3 in) tail[61]

Habitat: Forest[62]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown Population declining[62]

Dhofar shrew


C. dhofarensis
(Hutterer, 1988)
Yemen and Oman Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[63]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[63]

Doucet's musk shrew


C. douceti
Heim de Balsac, 1958
Western Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[64]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[64]

Dsinezumi shrew

Drawing of gray shrew

C. dsinezumi
(Temminck, 1842)
Japan
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland, grassland, and inland wetlands[65]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[65]

East African highland shrew


C. allex
Osgood, 1910
Kenya and Tanzania
Map of range
Size: 5–7 cm (2–3 in) long, plus 4–6 cm (2 in) tail[66]

Habitat: Grassland and inland wetlands[67]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[67]

Egyptian pygmy shrew


C. religiosa
(Geoffroy, 1827)
Northern Egypt
Map of range
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[68]

Habitat: [69]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[69]

Eisentraut's shrew


C. eisentrauti
Heim de Balsac, 1957
Cameroon
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland and forest[70]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown Population steady[70]

Elgon shrew


C. elgonius
Osgood, 1910
Kenya and Tanzania
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[71]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population declining[71]

Elongated shrew


C. elongata
Miller & Hollister, 1921
Sulawesi island in Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[72]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population declining[72]

Fingui white-toothed shrew


C. fingui
Ceríaco et al., 2015
Príncipe in São Tomé and Príncipe Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[73]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[73]

Fischer's shrew


C. fischeri
Pagenstecher, 1885
Kenya and Tanzania
Map of range
Size: 6–10 cm (2–4 in) long, plus 4–6 cm (2 in) tail[74]

Habitat: Savanna[75]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[75]

Flat-headed shrew


C. planiceps
Heller, 1910
West-central Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Inland wetlands[76]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[76]

Flower's shrew


C. floweri
Dollman, 1915
Northern Egypt
Map of range
Size: 5–8 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[77]

Habitat: Artificial terrain[78]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[78]

Fox's shrew


C. foxi
Dollman, 1915
Western and central Africa
Map of range
Size: 9–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[79]

Habitat: Savanna and forest[80]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[80]

Fraser's musk shrew


C. poensis
(Fraser, 1843)
Western Africa
Map of range
Size: 7–10 cm (3–4 in) long, plus 4–7 cm (2–3 in) tail[81]

Habitat: Shrubland and forest[82]

Diet: Arthropods and other invertebrates[81]
 LC 


Unknown Unknown[82]

Gathorne's shrew


C. gathornei
Jenkins, 2013
Himalayas in India Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and grassland[83]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[83]

Glass's shrew


C. glassi
Heim de Balsac, 1966
Ethiopia
Map of range
Size: 6–9 cm (2–4 in) long, plus 4–6 cm (2 in)tail[84]

Habitat: Shrubland, grassland, and inland wetlands[85]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown Population declining[85]

Gmelin's white-toothed shrew


C. gmelini
Pallas, 1811
Central Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland and grassland[86]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[86]

Goliath shrew


C. goliath
Thomas, 1906
Central Africa
Map of range
Size: 15–18 cm (6–7 in) long, plus 10–14 cm (4–6 in) tail[87]

Habitat: Forest[88]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[88]

Gracile naked-tailed shrew


C. maurisca
Thomas, 1904
Central Africa
Map of range
Size: 7–10 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[89]

Habitat: Inland wetlands and forest[90]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[90]

Grasse's shrew


C. grassei
Brosset, Dubost, & Heim de Balsac, 1965
West-central Africa
Map of range
Size: 7–9 cm (3–4 in) long, plus 6–8 cm (2–3 in) tail[91]

Habitat: Forest[92]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[92]

Greater Mindanao shrew


C. grandis
Miller, 1911
Philippines
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[93]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[93]

Greater red musk shrew

Drawing of brown shrew

C. flavescens
(Geoffroy, 1827)
Southern Africa
Map of range
Size: 8–12 cm (3–5 in) long, plus 3–6 cm (1–2 in) tail[94]

Habitat: Savanna and grassland[95]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[95]

Greater white-toothed shrew

Brown shrews

C. russula
(Hermann, 1780)
Europe and northwestern Africa
Map of range
Size: 6–9 cm (2–4 in) long, plus 3–5 cm (1–2 in) tail[96]

Habitat: Shrubland[97]

Diet: Invertebrates, as well as small mammals and small lizards[96]
 LC 


Unknown Population steady[97]

Greenwood's shrew


C. greenwoodi
Heim de Balsac, 1966
Somalia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, savanna, and shrubland[98]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[98]

Guramba shrew


C. phaeura
Osgood, 1936
Ethiopia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[99]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Unknown[99]

Guy's shrew


C. guy
Jenkins, Lunde, & Moncrieff, 2009
Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[100]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[100]

Güldenstädt's shrew


C. gueldenstaedtii
(Pallas, 1811)
Europe and western Asia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland, grassland, inland wetlands, rocky areas, intertidal marine, and coastal marine[101]

Diet: Insects, as well as molluscs, amphipods, annelids, and small vertebrates[101]
 LC 


Unknown Unknown[101]

Hainan Island shrew


C. wuchihensis
Sung, 1966
Southern China and northern Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[102]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[102]

Harenna shrew


C. harenna
Hutterer & Yalden, 1990
Ethiopia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[103]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Population declining[103]

Heather shrew


C. erica
Dollman, 1915
Angola
Map of range
Size: About 10 cm (4 in) long, plus 5 cm (2 in) tail[104]

Habitat: Unknown[105]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[105]

Hildegarde's shrew


C. hildegardeae
Thomas, 1904
Central Africa
Map of range
Size: 6–8 cm (2–3 in) long, plus 3–6 cm (1–2 in) tail[106]

Habitat: Forest[107]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[107]

Hill's shrew


C. hilliana
Jenkins & Smith, 1995
Thailand and Laos
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[108]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[108]

Horsfield's shrew


C. horsfieldii
(Tomes, 1856)
Scattered southern Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[109]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[109]

Hun shrew


C. attila
Dollman, 1915
Central Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[110]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[110]

Hutan shrew


C. hutanis
Ruedi & Vogel, 1995
Sumatra island in Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[111]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[111]

Indochinese shrew


C. indochinensis
Kloss, 1922
Southeastern Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[112]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[112]

Iranian shrew


C. susiana
Redding & Lay, 1978
Iran
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[113]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[113]

Jackass shrew


C. arispa
Spitzenberger, 1971
Turkey
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas[114]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[114]

Jackson's shrew


C. jacksoni
Thomas, 1904
East-central Africa
Map of range
Size: 5–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[115]

Habitat: Forest[116]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[116]

Javan ghost shrew


C. umbra
Demos, Achmadi, Handika, Maharadatunkamsi, Rowe, & Esselstyn, 2016
Java island in Indonesia Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[117]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown Unknown[117]

Javanese shrew


C. maxi
Sody, 1936
Indonesia and East Timor
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[118]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[118]

Jenkins's shrew


C. jenkinsi
Chakraborty, 1978
South Andaman Island in India
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[119]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Population declining[119]

Jouvenet's shrew


C. jouvenetae
Heim de Balsac, 1958
Western Africa
Map of range
Size: 6–9 cm (2–4 in) long, plus 4–7 cm (2–3 in) tail[120]

Habitat: Forest[121]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[121]

Kashmir white-toothed shrew


C. pullata
Miller, 1911
Northern India and Pakistan
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[122]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[122]

Katinka's shrew


C. katinka
Bate, 1937
Western Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[123]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[123]

Kilimanjaro shrew


C. monax
Thomas, 1910

Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[124]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[124]

Kinabalu shrew


C. baluensis
Thomas, 1898
Northern Borneo
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[125]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[125]

Kivu long-haired shrew


C. lanosa
Heim de Balsac, 1968
Central Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and inland wetlands[126]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown Population declining[126]

Kivu shrew


C. kivuana
Heim de Balsac, 1968
Central Africa
Map of range
Size: 6–9 cm (2–4 in) long, plus 6–8 cm (2–3 in) tail[127]

Habitat: Forest and inland wetlands[128]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown Unknown[128]

Lamotte's shrew


C. lamottei
Heim de Balsac, 1968
Western Africa
Map of range
Size: 8–9 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[127]

Habitat: Forest and savanna[129]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[129]

Large-headed forest shrew


C. grandiceps
Hutterer, 1983
Western Africa
Map of range
Size: 9–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[130]

Habitat: Forest[131]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[131]

Latona's shrew


C. latona
Hollister, 1916
Democratic Republic of the Congo
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[132]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[132]

Lesser Ryukyu shrew


C. watasei
Kuroda, 1924
Southern Japan
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland, grassland, and inland wetlands[133]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[133]

Lesser gray-brown musk shrew


C. silacea
Thomas, 1895
Southern Africa
Map of range
Size: 6–8 cm (2–3 in) long, plus 4–6 cm (2 in) tail[134]

Habitat: Rocky areas, grassland, shrubland, and savanna[135]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[135]

Lesser red musk shrew

Brown shrew

C. hirta
Peters, 1852
Southern and southeastern Africa
Map of range
Size: 6–10 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[136]

Habitat: Savanna, grassland, and desert[137]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[137]

Lesser rock shrew


C. serezkyensis
Laptev, 1929
Western Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Unknown[138]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[138]

Lesser white-toothed shrew

Gray shrew

C. suaveolens
(Pallas, 1811)
Europe and Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, grassland, rocky areas, and coastal/supratidal marine[139]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[139]

Long-footed shrew


C. crenata
Brosset, Dubost, & Heim de Balsac, 1965
West-central Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[140]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[140]

Long-tailed musk shrew


C. dolichura
Peters, 1876
Central Africa
Map of range
Size: 6–7 cm (2–3 in) long, plus 7–8 cm (3 in) tail[141]

Habitat: Forest[142]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[142]

Lucina's shrew


C. lucina
Dippenaar, 1980
Ethiopia
Map of range
Size: 8–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail[143]

Habitat: Grassland and inland wetlands[144]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown Population steady[144]

Ludia's shrew


C. ludia
Hollister, 1916
Central Africa
Map of range
Size: 6–7 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[145]

Habitat: Forest[146]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[146]

Luzon shrew


C. grayi
Dobson, 1890
Philippines
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[147]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[147]

Lwiro shrew


C. lwiroensis
Peterhans & Hutterer, 2013
Democratic Republic of the Congo Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[148]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[148]

MacArthur's shrew


C. macarthuri
St. Leger, 1934
Eastern Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[149]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[149]

MacMillan's shrew


C. macmillani
Dollman, 1915
Ethiopia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and savanna[150]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown Unknown[150]

Makwassie musk shrew


C. maquassiensis
Roberts, 1946
Southern Africa
Map of range
Size: 5–8 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[151]

Habitat: Grassland and rocky areas[152]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[152]

Malayan shrew


C. malayana
Robinson & Kloss, 1911
Malaysia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[153]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[153]

Mamfe shrew


C. virgata
Sanderson, 1940
West-central Africa Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[154]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[154]

Manenguba shrew


C. manengubae
Hutterer, 1982
West-central Africa
Map of range
Size: 7–9 cm (3–4 in) long, plus 6–7 cm (2–3 in) tail[151]

Habitat: Forest, shrubland, and grassland[155]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown Population declining[155]

Mauritanian shrew


C. lusitania
Dollman, 1915
Western and eastern Africa
Map of range
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[156]

Habitat: Savanna and shrubland[157]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[157]

Mduma's shrew


C. mdumai
Stanley, Hutterer, Giarla, & Esselstyn, 2015
Tanzania Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[158]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Unknown[158]

Mindanao shrew

Gray shrew

C. beatus
Miller, 1910
Philippines
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[159]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population declining[159]

Mindoro shrew


C. mindorus
Miller, 1910
Philippines
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[160]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[160]

Montane white-toothed shrew


C. montis
Thomas, 1906
Eastern Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, and grassland[161]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[161]

Moonshine shrew


C. luna
Dollman, 1910
Southeastern Africa
Map of range
Size: 6–11 cm (2–4 in) long, plus 3–7 cm (1–3 in) tail[156]

Habitat: Forest[162]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[162]

Mossy forest shrew


C. musseri
Ruedi & Vogel, 1995
Sulawesi island in Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[163]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[163]

Mount Nimba giant forest shrew


C. nimbasilvanus
Hutterer, 2003
Western Africa Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[164]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[164]

Munissi's shrew


C. munissii
Stanley, Hutterer, Giarla, & Esselstyn, 2015
Tanzania Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[165]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[165]

Narrow-headed shrew


C. stenocephala
Heim de Balsac & Dieterlen, 1979
Central Africa
Map of range
Size: 8–11 cm (3–4 in) long, plus 6–9 cm (2–4 in) tail[166]

Habitat: Inland wetlands[167]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown Population declining[167]

Negev shrew


C. ramona
Ivanitskaya, Shenbrot, & Nevo, 1996
Israel
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Desert[168]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[168]

Negros shrew


C. negrina
Rabor, 1952
Philippines
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[169]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[169]

Nicobar shrew


C. nicobarica
Miller, 1902
Great Nicobar Island in India
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[170]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Population declining[170]

Nigerian shrew


C. nigeriae
Dollman, 1915
West-central Africa
Map of range
Size: 9–11 cm (4 in) long, plus 5–7 cm (2–3 in) tail[171]

Habitat: Forest[172]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[172]

Nimba shrew


C. nimbae
Heim de Balsac, 1956
Western Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[173]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown Population declining[173]

Niobe's shrew


C. niobe
Thomas, 1906
Central Africa
Map of range
Size: 6–9 cm (2–4 in) long, plus 5–7 cm (2–3 in) tail[174]

Habitat: Forest and grassland[175]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[175]

North African white-toothed shrew

Brown shrew

C. pachyura
Küster, 1835
Northern Africa and southern Europe
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland[176]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[176]

Nyiro shrew


C. macowi
Dollman, 1915
Kenya
Map of range
Size: 6–8 cm (2–3 in) long, plus 5–6 cm (2–2 in) tail[177]

Habitat: Unknown[178]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[178]

Oriental shrew

Brown shrew

C. orientalis
Jentink, 1890

Two subspecies
  • C. o. lawuana
  • C. o. orientalis
Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[179]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[179]

Palawan shrew


C. palawanensis
Taylor, 1934
Philippines
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[180]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[180]

Pale gray shrew


C. pergrisea
Miller, 1913
Pakistan
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[181]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[181]

Panay shrew


C. panayensis
Hutterer, 2007
Philippines
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[182]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[182]

Peninsular shrew


C. negligens
Kloss, 1914
Southeastern Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[183]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[183]

Peters's musk shrew


C. gracilipes
Peters, 1870
Tanzania
Map of range
Size: About 6 cm (2 in) long, plus 5 cm (2 in) tail[87]

Habitat: Unknown[184]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[184]

Phan Luong's shrew


C. phanluongi
Jenkins, Abramov, Rozhnov, & Olsson, 2010
Vietnam and Cambodia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[185]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[185]

Phu Hoc shrew


C. phuquocensis
Abramov, Jenkins, Rozhnov, & Kalinin, 2008
Phú Quốc island in Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[186]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[186]

Pitman's shrew


C. pitmani
Barclay, 1932
Zambia
Map of range
Size: About 5 cm (2 in) long, plus 4 cm (2 in) tail[187]

Habitat: Unknown[188]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[188]

Polia's shrew


C. polia
Hollister, 1916
Democratic Republic of the Congo
Map of range
Size: About 6 cm (2 in) long, plus 7 cm (3 in) tail[81]

Habitat: Unknown[189]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[189]

Rainey's shrew


C. raineyi
Heller, 1912
Kenya
Map of range
Size: 9–10 cm (4 in) long, plus 5–7 cm (2–3 in) tail[190]

Habitat: Forest[191]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[191]

Reddish-gray musk shrew


C. cyanea
(Duvernoy, 1838)
Southern Africa
Map of range
Size: 6–10 cm (2–4 in) long, plus 3–7 cm (1–3 in) tail[192]

Habitat: Forest, grassland, and caves[193]

Diet: Invertebrates[193]
 LC 


Unknown Population steady[193]

Roosevelt's shrew


C. roosevelti
(Heller, 1910)
Central Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[194]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[194]

Ryukyu shrew


C. orii
Kuroda, 1924
Southern Japan
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[195]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[195]

Sa Pa shrew

C. sapaensis
Jenkins, Abramov, Bannikova, & Rozhnov, 2013
Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[196]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[196]

Saharan shrew


C. tarfayensis
Vesmanis & Vesmanis, 1980
Mauritania and Morocco
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Rocky areas and desert[197]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Population declining[197]

Sahelian tiny shrew


C. pasha
Dollman, 1915
Scattered northern Africa
Map of range
Size: 4–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail[198]

Habitat: Savanna[199]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[199]

Savanna dwarf shrew


C. nanilla
Thomas, 1909
Western and eastern Africa
Map of range
Size: 4–6 cm (2–2 in) long, plus 3–5 cm (1–2 in) tail[200]

Habitat: Savanna[201]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[201]

Savanna path shrew


C. viaria
(Geoffroy, 1834)
Western, central, and eastern Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and savanna[202]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[202]

Savanna shrew


C. fulvastra
(Sundevall, 1843)
Central Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[203]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[203]

Savanna swamp shrew


C. longipes
Hutterer & Happold, 1983
West-central Africa
Map of range
Size: 9–12 cm (4–5 in) long, plus 5–7 cm (2–3 in) tail[204]

Habitat: Inland wetlands[205]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[205]

Siberian shrew


C. sibirica
Dukelsky, 1930
Central Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, grassland, inland wetlands, and desert[206]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[206]

Sibuyan shrew


C. ninoyi
Esselstyn & Goodman, 2010
Philippines
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[207]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[207]

Sicilian shrew

Brown shrew

C. sicula
Miller, 1900

Four subspecies
  • C. s. aegatensis
  • C. s. calypso
  • C. s. esuae
  • C. s. sicula
Italy and Malta
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Shrubland[208]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[208]

Sinharaja white-toothed shrew


C. hikmiya
Meegaskumbura, Meegaskumbura, Pethiyagoda, Manamendra-Arachchi, & Schneider, 2007
Sri Lanka
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[209]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[209]

Small-footed shrew


C. parvipes
Osgood, 1910
Central Africa
Map of range
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[210]

Habitat: Forest, savanna, and shrubland[211]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[211]

Smoky white-toothed shrew


C. fumosa
Thomas, 1904
Kenya
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[212]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population declining[212]

Sokolov's shrew


C. sokolovi
Jenkins, Abramov, Rozhnov, & Makarova, 2007
Vietnam Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[213]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[213]

Somali dwarf shrew


C. nana
Dobson, 1890
Ethiopia and Somalia
Map of range
Size: About 4 cm (2 in) long, plus 3 cm (1 in) tail[214]

Habitat: Grassland[215]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[215]

Somali shrew


C. somalica
Thomas, 1895
Eastern Africa
Map of range
Size: 6–8 cm (2–3 in) long, plus 4–5 cm (2–2 in) tail[166]

Habitat: Savanna[216]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[216]

Southeast Asian shrew

Gray shrew

C. fuliginosa
(Blyth, 1856)
Southeastern Asian
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest, shrubland, grassland, and rocky areas[217]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[217]

Sri Lankan long-tailed shrew


C. miya
Phillips, 1929
Sri Lanka
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[218]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[218]

Sulawesi shrew


C. lea
Miller & Hollister, 1921
Sulawesi island in Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[219]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[219]

Sulawesi tiny shrew


C. levicula
Miller & Hollister, 1921
Sulawesi island in Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[220]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[220]

Sulawesi white-handed shrew


C. rhoditis
Miller & Hollister, 1921
Sulawesi island in Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[221]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[221]

Sumatran giant shrew


C. lepidura
Lyon, 1908
Sumatra island in Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and shrubland[222]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[222]

Sumatran long-tailed shrew


C. paradoxura
Dobson, 1886
Sumatra island in Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[223]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population declining[223]

Sunda shrew

C. monticola
Peters, 1870
Southeastern Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[224]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[224]

Swamp musk shrew


C. mariquensis
(Smith, 1844)

Three subspecies
  • C. m. mariquensis
  • C. m. neavei
  • C. m. shortridgei
Southern Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Inland wetlands[225]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[225]

São Tomé shrew

Brown shrew

C. thomensis
(Bocage, 1887)
São Tomé Island in São Tomé and Príncipe
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[226]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[226]

Taiwanese gray shrew


C. tanakae
Kuroda, 1938
Eastern Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest and grassland[227]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[227]

Tanzanian shrew


C. tansaniana
Hutterer, 1986
Tanzania
Map of range
Size: 8–11 cm (3–4 in) long, plus 5–8 cm (2–3 in) tail[228]

Habitat: Forest[229]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 NT 


Unknown Unknown[229]

Tarella shrew


C. tarella
Dollman, 1915
Central Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[230]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 EN 


Unknown Population declining[230]

Telford's shrew


C. telfordi
Hutterer, 1986
Tanzania
Map of range
Size: 10–11 cm (4 in) long, plus 7–9 cm (3–4 in) tail[231]

Habitat: Forest[232]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown Population declining[232]

Thalia's shrew


C. thalia
Dippenaar, 1980
Ethiopia
Map of range
Size: 7–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[231]

Habitat: Forest, savanna, shrubland, grassland, and inland wetlands[233]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[233]

Therese's shrew


C. theresae
Heim de Balsac, 1968
Western Africa
Map of range
Size: 7–11 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[234]

Habitat: Forest and savanna[235]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[235]

Thick-tailed shrew


C. brunnea
Jentink, 1888

Two subspecies
  • C. b. brunnea
  • C. b. pudjonica
Indonesia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[236]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[236]

Timor shrew


C. tenuis
(Müller, 1840)
Timor
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[237]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[237]

Turbo shrew


C. turba
Dollman, 1910
Central Africa
Map of range
Size: 9–11 cm (4 in) long, plus 5–6 cm (2–2 in) tail[238]

Habitat: Forest[239]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[239]

Ugandan lowland shrew


C. selina
(Dollman, 1915)
Kenya and Uganda
Map of range
Size: 7–9 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[240]

Habitat: Forest and inland wetlands[241]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[241]

Ugandan musk shrew


C. mutesae
Heller, 1910
Central Africa
Map of range
Size: About 11 cm (4 in) long, plus 6 cm (2 in) tail[214]

Habitat: Forest[242]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[242]

Ultimate shrew


C. ultima
Dollman, 1915
Kenya
Map of range
Size: About 9 cm (4 in) long, plus 6 cm (2 in) tail[243]

Habitat: Forest[244]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[244]

Upemba shrew


C. zimmeri
Osgood, 1936
Democratic Republic of the Congo
Map of range
Size: 11–13 cm (4–5 in) long, plus 4–6 cm (2–2 in) tail[245]

Habitat: Unknown[246]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[246]

Usambara shrew


C. usambarae
Dippenaar, 1980
Tanzania
Map of range
Size: 7–9 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[247]

Habitat: Forest[248]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 VU 


Unknown Unknown[248]

Ussuri white-toothed shrew

Brown shrew

C. lasiura
(Dobson, 1890)
Eastern Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Grassland[249]

Diet: Insects and other invertebrates, as well as small vertebrates[249]
 LC 


Unknown Unknown[249]

Voi shrew


C. voi
Osgood, 1910
Central and eastern Africa
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Savanna[250]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[250]

Voracious shrew

Brown shrew

C. vorax
Allen, 1923
Southeastern Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Forest[251]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[251]

West African long-tailed shrew


C. muricauda
(Miller, 1900)
Western Africa
Map of range
Size: 5–7 cm (2–3 in) long, plus tail[252]

Habitat: Forest[253]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[253]

West African pygmy shrew


C. obscurior
Heim de Balsac, 1958
Western Africa
Map of range
Size: 4–5 cm (2–2 in) long, plus 3–4 cm (1–2 in) tail[254]

Habitat: Forest[255]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[255]

Whitaker's shrew


C. whitakeri
De Winton, 1898
Northwestern Africa
Map of range
Size: 5–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[256]

Habitat: Coastal marine, intertidal marine, and shrubland[257]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[257]

Wimmer's shrew


C. wimmeri
Heim de Balsac & Aellen, 1958
Ivory Coast
Map of range
Size: 8–9 cm (3–4 in) long, plus 5–6 cm (2–2 in) tail[258]

Habitat: Forest[259]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 CR 


Unknown Unknown[259]

Xanthippe's shrew


C. xantippe
Osgood, 1910
Kenya and Tanzania
Map of range
Size: 8–10 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail[260]

Habitat: Savanna and shrubland[261]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[261]

Yankari shrew


C. yankariensis
Hutterer & Jenkins, 1980
Central and eastern Africa
Map of range
Size: 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[260]

Habitat: Savanna[262]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Population steady[262]

Zaphir's shrew


C. zaphiri
Dollman, 1915
Ethiopia and Kenya
Map of range
Size: About 10 cm (4 in) long, plus 6 cm (2 in) tail[263]

Habitat: Unknown[264]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 DD 


Unknown Unknown[264]

Zarudny's rock shrew

Gray shrew

C. zarudnyi
Ognew, 1928
Central Asia
Map of range
Size: 4–18 cm (2–7 in) long, plus 4–11 cm (2–4 in) tail[6]

Habitat: Desert[265]

Diet: Invertebrates, small mammals, frogs, toads, and lizards[6]
 LC 


Unknown Unknown[265]

Genus Diplomesodon Brandt, 1852 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Piebald shrew

Gray and white shrew

D. pulchellum
(Lichtenstein, 1823)
Central Asia
Map of range
Size: 5–8 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[266]

Habitat: Desert[267]

Diet: Insects, as well as small lizards[267]
 LC 


Unknown Population steady[267]

Genus Feroculus Kelaart, 1852 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Kelaart's long-clawed shrew


F. feroculus
(Kelaart, 1850)
Southern India and Sri Lanka
Map of range
Size: 10–12 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[268]

Habitat: Forest, grassland, and inland wetlands[269]

Diet: Earthworms, small vertebrates, and plants