2016 in paleontology

From Wikipedia the free encyclopedia

List of years in paleontology (table)
In paleobotany
2013
2014
2015
2016
2017
2018
2019
In arthropod paleontology
2013
2014
2015
2016
2017
2018
2019
In paleoentomology
2013
2014
2015
2016
2017
2018
2019
In paleomalacology
2013
2014
2015
2016
2017
2018
2019
In paleoichthyology
2013
2014
2015
2016
2017
2018
2019
In reptile paleontology
2013
2014
2015
2016
2017
2018
2019
In archosaur paleontology
2013
2014
2015
2016
2017
2018
2019
In mammal paleontology
2013
2014
2015
2016
2017
2018
2019

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2016.

Flora[edit]

Plants[edit]

Fungi[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Cultoraquaticus[2]

Gen. et sp. nov

Valid

Strullu-Derrien in Strullu-Derrien et al.

Early Devonian

Rhynie chert

 United Kingdom

A fungus belonging to the group Chytridiomycetes.
The type species is C. trewini.

Dennisiellinites[3]

Gen. et sp. nov

Valid

Bannister, Conran & Lee

Eocene

Pikopiko Fossil Forest

 New Zealand

A fungus. Genus includes new species D. pikopikoensis .

Gerontomyces[4]

Gen. et sp. nov

Valid

Poinar

Eocene

Baltic amber

Europe (Baltic Sea coast)

A gilled mushroom. The type species is G. lepidotus.

Hypoxylonites foyelensis[5]

Sp. nov

Valid

Bianchinotti, Martínez & Saxena in Martínez et al.

Paleogene (probably Eocene)

Troncoso Formation

 Argentina

A fungus described on the basis of spores.

Monodictysporites[6]

Gen. et sp. nov

Valid

Klymiuk

Ypresian

Allenby Formation
Princeton Chert

 Canada
( British Columbia)

A ascomycotan fungal spore taxon.
The type species is M. princetonensis.
Likely hosted on Dennstaedtiopsis aerenchymata

Paleocadus[7]

Gen. et sp. nov

Valid

Poinar

Cretaceous

Burmese amber

 Myanmar

An Eccrinaceous fungus. Type species P. burmiticus.

Pluricellaesporites patagonicus[5]

Sp. nov

Valid

Bianchinotti, Martínez & Saxena in Martínez et al.

Paleogene

 Argentina

A fungus described on the basis of spores resembling those of members of the genus Bactrodesmium.

Trewinomyces[8]

Gen. et sp. nov

Valid

Krings, Taylor & Martin

Early Devonian

Rhynie chert

 United Kingdom

A fungus of uncertain phylogenetic placement,
showing similarities to Macrochytrium (Chytridiomycota) and Blastocladiella (Blastocladiomycota) species.
The type species is T. annulifer.

Cnidarians[edit]

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Aphyllum pachystele[11]

Sp. nov

Valid

Munson & Jell

Silurian (late Wenlock)

Jack Formation

 Australia

A rugose coral belonging to the family Tryplasmatidae.

Aulokoninckophyllum potyi[12]

Sp. nov

Valid

Somerville, Rodríguez & Said

Carboniferous

Azrou-Khenifra Basin

 Morocco

An aulate coral.

Axuolites higoensis[13]

Sp. nov

Valid

Niko

Silurian (Ludlow)

Fukami Formation

 Japan

A tabulate coral belonging to the order Favositida and the family Coenitidae.

Bjarmia[14]

Gen. et sp. nov

Valid

Grazhdankin

Late Ediacaran (~550 Ma)

Erga Formation

 Russia

A jellyfish-like organism, a possible relative of scyphomedusae. Genus includes new species B. cycloplerusa.

Byronia jaegeri[15]

Sp. nov

Valid

Vinn et al.

Ordovician (Katian)

 Estonia

A possible thecate scyphozoan.

Calceola collini[16]

Sp. nov

Valid

Wright, Plusquellec & Gourvennec

Devonian (Eifelian)

Saint-Fiacre Formation

 France

An operculate coral belonging to the family Calceolidae; a species of Calceola.

Cambroctoconus coreaensis[17]

Sp. nov

Valid

Park et al.

Cambrian

Daegi Formation

 South Korea

A stem-cnidarian.

Caninophyllum charli[18]

Sp. nov

Valid

Denayer

Carboniferous (Tournaisian)

Yılanlı Formation

 Turkey

A rugose coral belonging to the family Bothrophyllidae; a species of Caninophyllum.

Cystodactylon[19]

Gen. et sp. nov

Valid

Denayer & Webb

Carboniferous (Tournaisian)

Gudman Formation

 Australia

A rugose coral. Genus includes new species C. orbum.

Dokophyllum hillae[11]

Sp. nov

Valid

Munson & Jell

Silurian (Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Ketophyllidae.

Edaphophyllum zhaotongense[20]

Sp. nov

Valid

Liao & Ma

Devonian (Emsian)

 China

A rugose coral.

Eoaequorea[21]

Gen. et sp. nov

Valid

Tang et al.

Ediacaran

Doushantuo Formation

 China

A member of Hydrozoa of uncertain phylogenetic placement. The type species is Eoaequorea xingi.

Eosiderastrea[22]

Gen. et comb. et 2 sp. nov

Valid

Löser

Cretaceous (late Valanginian to Santonian)

 Austria
 France
 Germany
 Greece
 Iran
 Italy
 Japan
 Mexico
 Poland
 Spain
 Ukraine
 United States
 Uzbekistan

A stony coral belonging to the family Siderastreidae. The type species is "Siderastrea" cuyleri Wells (1932); genus also includes "Phyllocoenia" cotteaui Orbigny (1850), "Diploastrea" crassa Kuzmicheva (1980), "Diploastrea" crassicostata Morycowa & Masse (1998), "Phyllocoenia" cyclops Felix (1891), "Confusastrea" dollfusi Prever (1909), "Confusastrea" felixi Prever (1909); "Stephanocoenia" grandipora Orbigny (1849), "Diploastrea" harrisi Wells (1932), "Diploastrea" hilli Wells (1933), "Montastraea" nagaoi Eguchi (1951), "Pleurocora" reussi Milne Edwards (1857), "Diplocoenia" splendida Prever (1909), "Diploastrea" vaughani Wells (1933) and "Plesiofavia" villaltai Reig Oriol (1991), as well as new species E. paragrandipora Löser (2016) and E. stefani Löser (2016).[23]

Famastraea[24]

Gen. et sp. nov

Valid

Berkowski, Zapalski & Wrzołek

Devonian (Famennian)

 Poland

A coral belonging to the group Rugosa and the family Campophyllidae. The type species is Famastraea catenata.

Fomichevella najafi[25]

Sp. nov

Valid

Badpa et al.

Carboniferous (Bashkirian)

Sardar Formation

 Iran

A member of Rugosa belonging to the group Stauriida and the family Kleopatrinidae.

Gertholites haikawai[26]

Sp. nov

Valid

Niko & Fujikawa

Early Carboniferous

Akiyoshi Limestone Group

 Japan

A tabulate coral.

Gerviphyllum[16]

Gen. et comb. nov

Valid

Wright, Plusquellec & Gourvennec

Early Devonian (likely late Lochkovian to early Pragian)

 France

An operculate coral belonging to the family Calceolidae; a new genus for "Calceola" gervillei Bayle (1878).

Gudmania[19]

Gen. et sp. nov

Valid

Denayer & Webb

Carboniferous (Tournaisian)

Gudman Formation

 Australia

A rugose coral. Genus includes new species G. darumbalae.

Heintzella fluegeli[25]

Sp. nov

Valid

Badpa et al.

Carboniferous (Bashkirian)

Sardar Formation

 Iran

A member of Rugosa belonging to the group Stauriida and the family Kleopatrinidae.

Kullingia rotadiscopsis[21]

Sp. nov

Valid

Tang et al.

Ediacaran

Doushantuo Formation

 China

A member of Hydrozoa of uncertain phylogenetic placement; a species of Kullingia.

Madrepora mitsukurii[27]

Sp. nov

Valid

Niko, Suzuki & Taguchi

Miocene

Katsuta Group

 Japan

A madrepore.

Marekostragulum grafi[28]

Sp. nov

Valid

Plusquellec, Galle & Franke

Devonian (Emsian)

 Germany

A tabulate coral belonging to the family Hyostragulidae.

Mesoconularia (?) lukesi[29]

Sp. nov

Valid

Mergl, Ferrova & Frýda

Devonian (late Emsian)

Suchomasty Limestone

 Czech Republic

A member of Conulariida, possibly a species of Mesoconularia.

?Michelinia vinni[30]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Micheliniidae; possibly a species of Michelinia.

Multicarinophyllum vepreculatum[11]

Sp. nov

Valid

Munson & Jell

Silurian (Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Amsdenoididae.

Oligophylloides maroccanus[31]

Sp. nov

Valid

Weyer

Devonian (late Famennian)

 Morocco

A member of Heterocorallia.

Palaeosiderofungia[22]

Gen. et comb. nov

Valid

Löser

Cretaceous (Aptian to Santonian)

 Austria
 Czech Republic
 France
 Germany
 Greece
 Serbia

A stony coral belonging to the family Siderastreidae; a new genus for "Thamnasteria" exigua Reuss (1854).

Parostragulum[28]

Gen. et sp. nov

Valid

Plusquellec, Galle & Franke

Devonian (Emsian)

 Germany

A tabulate coral belonging to the family Hyostragulidae. The type species is P. problematicum.

Pleurodictyum goldfussi[32]

Sp. nov

Valid

Plusquellec & Franke

Devonian (Emsian)

Wiltz Beds

 Germany
 Luxembourg

A tabulate coral belonging to group Favositida and to the family Micheliniidae.

Pleurodictyum pruemensis[32]

Sp. nov

Valid

Plusquellec & Franke

Devonian (Emsian)

Wiltz Beds

 Germany

A tabulate coral belonging to group Favositida and to the family Micheliniidae.

Potyphyllum[33]

Gen. et comb. nov

Valid

Coen-Aubert

Devonian (late Frasnian)

 Belgium
 Czech Republic
 France
 Germany
 Poland
 Russia
 United Kingdom

A rugose coral belonging to the family Phillipsastreidae. The type species is "Cyathophyllum" ananas Goldfuss (1826); genus also includes "Pseudoacervularia" dybowskii Różkowska (1953), "Pseudoacervularia" intercellulosa (Phillips, 1841) sensu Pickett (1967), "Schlüteria" lyskovensis Ermakova (1957), "Acervularia" macrommata Roemer (1855), "Phillipsastrea" plantana Różkowska (1979), "Cyathophyllum" profundum Michelin (1845), "Phillipsastrea" rozkowskae Scrutton (1968), "Pseudoacervularia" cf. smithi (Różkowska, 1953) sensu Pickett (1967), "Phillipsastrea" ananas veserensis Coen-Aubert (1974) and "Phillipsastrea" zerda Galle, 1992 in Hladil et al. (1992).

Preisingerella[34]

Gen. et sp. nov

Valid

Kossovaya, Novak & Weyer

Early Permian

 Slovenia

A coral similar to members of the genus Caninia. The type species is P. stegovnikensis.

Ptychophyllum variatum[11]

Sp. nov

Valid

Munson & Jell

Silurian (late Gorstian to Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Ptychophyllidae.

Pycnostylus polyphyllodus[11]

Sp. nov

Valid

Munson & Jell

Silurian (late Wenlock)

Jack Formation

 Australia

A rugose coral belonging to the family Pycnostylidae.

Sokolovia[35]

Gen. et sp. nov

Junior homonym

Tsyganko

Late Devonian (Famennian)

 Russia
( Komi Republic)

A tabulate coral belonging to the family Pachyporidae. The type species is Sokolovia pershinae. The generic name is preoccupied by Sokolovia Ilovaisky (1934) and Sokolovia Shishkinskaya (1964).

Spinaxon[36]

Gen. et sp. nov

Valid

Weyer

Devonian (late Frasnian)

 France
 Germany

A member of Rugosa belonging to the family Petraiidae. The type species is Spinaxon potyi.

Syringopora hilarowiczi[30]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Syringoporidae; a species of Syringopora.

Syringopora kowalensis[30]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Syringoporidae; a species of Syringopora.

Talfania[37]

Gen. et sp. nov

Valid

Peel & McDermott

Ordovician (Katian)

Sholeshook Limestone Formation

 United Kingdom

A solitary coral. The type species is Talfania calicula.

Thamnoptychia mistiaeni[30]

Sp. nov

Valid

Zapalski, Berkowski & Wrzołek

Devonian (late Famennian)

 Poland

A tabulate coral belonging to the family Pachyporidae; a species of Thamnoptychia.

Turbinolia (Turbinolia) deformis[38]

Sp. nov

Valid

Berezovsky in Berezovsky & Berezovsky

Eocene

 Ukraine

A stony coral.

Turbinolia (Turbinolia) emendata[38]

Sp. nov

Valid

Berezovsky in Berezovsky & Berezovsky

Eocene

 Ukraine

A stony coral.

Utaratuia yunnanensis[20]

Sp. nov

Valid

Liao & Ma

Devonian (Eifelian)

 China

A rugose coral.

Vassiljukia[39]

Gen. et comb. nov

Valid

Denayer & Ogar

Carboniferous (Viséan)

 Turkey
 Ukraine

A colonial rugose coral; a new genus for "Lithostrotion" columnariformis Vassiljuk (1960).

Vesicospina[11]

Gen. et sp. nov

Valid

Munson & Jell

Silurian (Ludfordian)

Jack Formation

 Australia

A rugose coral belonging to the family Kyphophyllidae. The type species is V. julli.

Vulykhia[40]

Nom. nov

Valid

Doweld

Late Ordovician

 Mongolia

An anthozoan belonging to the superfamily Heliolitoidea; a replacement name for Concavites Bondarenko & Minzhin (1981).

Arthropods[edit]

Bryozoans[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Anisotrypa kjarkiensis[41]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Armenia

A bryozoan belonging to the group Trepostomata and the family Anisotrypidae.

Beisselina skyscanica[42]

Sp. nov

Valid

Koromyslova & Pakhnevich

Late Cretaceous (Campanian)

 Belarus

A bryozoan belonging to the group Ascophora and the family Tessaradomidae; a species of Beisselina.

Celleporaria pirabasensis[43]

Sp. nov

Valid

Muricy et al.

Oligocene-Miocene

Pirabas Formation

 Brazil

A bryozoan belonging to the group Cheilostomata and the family Lepraliellidae.

Celleporaria triangulavicularis[43]

Sp. nov

Valid

Muricy et al.

Oligocene-Miocene

Pirabas Formation

 Brazil

A bryozoan belonging to the group Cheilostomata and the family Lepraliellidae.

Dybowskiella hupehensiformis[44]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Cystoporata belonging to the family Fistuliporidae.

Dyscritella lii[44]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Trepostomata belonging to the family Dystritellidae.

Ehrhardina[45]

Gen. et 2 sp. nov

Valid[46]

Martha & Taylor

Late Cretaceous (Cenomanian)

 France
 Germany
 United Kingdom

A bryozoan belonging to the group Flustrina and the family Onychocellidae. The type species is Ehrhardina voigti; genus also includes Ehrhardina pikeae.

Eridotrypella danzikensis[47]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata and the family Eridotrypellidae.

Etherella tibetensis[44]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Cystoporata belonging to the family Etherellidae.

Fistuliphragma moniliformis[48]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistuliphragma sibirica[48]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistulipora sakagamii[44]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China
 Thailand

A member of Cystoporata belonging to the family Fistuliporidae.

Fistulipora salairiensis[48]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistuliramus fasciculus[48]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Fistulocladia cincinnata[48]

Sp. nov

Valid

Mesentseva

Devonian (Emsian)

 Russia

A bryozoan belonging to the group Cystoporida.

Margaretta amplipora[49]

Sp. nov

Valid

Sonar & Gaikwad

Cenozoic

 India

An ascophoran belonging to the family Margarettidae.

Margaretta guhai[49]

Nom. nov

Valid

Sonar & Gaikwad

Cenozoic

 India

An ascophoran belonging to the family Margarettidae.

Margaretta hariparensis[49]

Sp. nov

Valid

Sonar & Gaikwad

Cenozoic

 India

An ascophoran belonging to the family Margarettidae.

Metastenodiscus[50]

Gen. et comb. nov

Valid

Ernst, Schäfer & Grant-Mackie

Late Triassic

 New Caledonia
 New Zealand

A trepostome bryozoan. A new genus for "Stenodiscus" zealandicus Schäfer & Grant-Mackie (1994) and "Stenodiscus" kawhiae Schäfer & Grant-Mackie (1994).

Nikiforopora arpaensis[41]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata and the family Stenoporidae.

Pachydermopora grodnoensis[42]

Sp. nov

Valid

Koromyslova & Pakhnevich

Late Cretaceous (Campanian)

 Belarus

A bryozoan belonging to the group Ascophora and the family Tessaradomidae; a species of Pachydermopora.

Petalotrypa myunkhbalaensis[47]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata.

Planicellaria walsariensis[51]

Sp. nov

Valid

Sonar & Pawar

Cenozoic

 India

A bryozoan belonging to the group Cheilostomata and the family Calloporidae.

Protoretepora irregularis[44]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Fenestrata belonging to the family Polyporidae.

Pyriporella charopadiensis[51]

Sp. nov

Valid

Sonar & Pawar

Cenozoic

 India

A bryozoan belonging to the group Cheilostomata and the family Calloporidae.

Pyriporella vadsariensis[51]

Sp. nov

Valid

Sonar & Pawar

Cenozoic

 India

A bryozoan belonging to the group Cheilostomata and the family Calloporidae.

Rectifenestella famenniensis[47]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Azerbaijan

A bryozoan belonging to the family Fenestellidae.

Rectifenestella kadrluiensis[47]

Sp. nov

Valid

Tolokonnikova

Devonian (Famennian)

 Armenia

A bryozoan belonging to the family Fenestellidae.

Stenophragmidium buckhornensis[52]

Sp. nov

Valid

Ernst et al.

Carboniferous (Pennsylvanian)

Boggy Formation

 United States
( Oklahoma)

Streblotrypa (Streblotrypa) heltzelae[52]

Sp. nov

Valid

Ernst et al.

Carboniferous (Pennsylvanian)

Boggy Formation

 United States
( Oklahoma)

Streblotrypa (Streblotrypa) parviformis[44]

Sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Cryptostomata belonging to the family Hyphasmoporidae.

Tabuliporella nakhichevanica[41]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Azerbaijan

A bryozoan belonging to the group Trepostomata and the family Crustoporidae.

Tibetiporella[44]

Gen. et sp. nov

Valid

Ernst

Permian

Zongba Formation

 China

A member of Fenestrata belonging to the family Polyporidae. The type species is T. ornata.

Timanotrypa australis[44]

Sp. nov

Valid

Ernst

Permian

Noonkanbah Formation
Rat Buri Limestone
Zongba Formation

 Australia
 China
 Thailand

A member of Cryptostomata belonging to the family Timanodictyidae.

Zigzagopora[53]

Gen. et sp. nov

Valid

Wilson & Taylor

Ordovician (Sandbian)

Bromide Formation

 United States
( Oklahoma)

A cyclostome bryozoan. Genus includes new species Z. wigleyensis.

Brachiopods[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Acritosia ogamensis[54]

Sp. nov

Valid

Tazawa et al.

Early Permian (Kungurian)

Nabeyama Formation

 Japan

Acrosaccus scutatus[55]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Discinidae.

Anathyris (Anathyris) calestiennensis[56]

Sp. nov

Valid

Mottequin et al.

Devonian (Frasnian)

Nismes Formation

 Belgium

Angulispirifer[57]

Gen. et comb. nov

Junior synonym

Waterhouse

Permian

 Australia

A member of Spiriferida. Genus includes "Spirifer" phalaena Dana (1849). Lee et al. (2023) subsequently considered the genus Angulispirifer to be a junior synonym of the genus Sulciplica.[58]

Anisopleurella antiqua[59]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Sowerbyellidae.

Apatobolus anoskelidion[55]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Obolidae.

Atansoria australis[55]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Obolidae.

Atryparia (Costatrypa) agricolae[60]

Sp. nov

Valid

Halamski & Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Atrypidae.

Aulacothyris maendlii[61]

Sp. nov

Valid

Sulser

Middle Jurassic (Callovian)

  Switzerland

A member of Terebratulida belonging to the family Zeilleriidae.

Aulacothyris waikatoensis[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Bellimurina fluctuosa[59]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Strophomenidae.

Biernatia pseudoplana[55]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Biernatiidae.

Biernatia wrighti[55]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Biernatiidae.

Biernatium minus[60]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Mystrophoridae.

Buxtonia inexpletucosta[63]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Canalilatus musculosus[64]

Sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A lingulate brachiopod belonging to the family Zhanatellidae.

Centreplicatus[65]

Gen. nov

Valid

Zeng,Zhang & Han in Zeng et al.

Ordovician (Hirnantian)

 China

A member of Pentamerida belonging to the family Pentameridae.

Cerasinella[66]

Nom. nov

Valid

Copper

Silurian (Llandovery)

Merrimack Formation

 Canada
( Quebec)

An atrypoid brachiopod; a replacement name for Cerasina Copper (1995) (preoccupied).

Chapinella belkovskensis[67]

Sp. nov

Valid

Baranov, Sokiran & Blodgett

Devonian (Famennian)

Nerpalakhsk Formation

 Russia
( Sakha Republic)

A member of Rhynchonellida belonging to the family Pugnacidae.

Chilcatreta[68]

Gen. et sp. nov

Valid

Lavié & Benedetto

Ordovician (Darriwilian)

San Juan Formation

 Argentina

A siphonotretid brachiopod. Genus includes new species C. tubulata.

Crispithyris[62]

Gen. et sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida. Genus includes new species C. nauarchus.

Cyrtina transcaucasica[69]

Sp. nov

Valid

Gretchishnikova & Oleneva in Oleneva

Devonian (Givetian)

 Azerbaijan

A member of Spiriferinida belonging to the family Cyrtinidae.

Davidsonia enmerkaris[60]

Sp. nov

Valid

Halamski in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Davidsoniidae.

Dictyoclostus transversum[63]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Disculina mancenidoi[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Dolerorthis nadruvensis[70]

Sp. nov

Valid

Paškevičius & Hints

Ordovician (Katian)

 Lithuania

A member of Orthida belonging to the family Hesperorthidae.

Dulankarella hyrcanica[59]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Leptellinidae.

Dyoros (Lissosia) maya[71]

Sp. nov

Valid

Torres-Martínez, Sour-Tovar & Barragán

Permian (Leonardian)

Paso Hondo Formation

 Mexico

Echinocoelia parva[60]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Ambocoeliidae.

Eochonetes maearum[72]

Sp. nov

Valid

Bauer & Stigall

Late Ordovician

 United States
( Wyoming)

Eochonetes minerva[72]

Sp. nov

Valid

Bauer & Stigall

Late Ordovician

 United States
( Texas)

Eochonetes voldemortus[72]

Sp. nov

Valid

Bauer & Stigall

Late Ordovician

Saturday Mountain Formation

 United States
( Idaho)

Eoconulus puteus[55]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Eoconulidae.

Eolingularia[73]

Gen. et comb. nov

Valid

Bitner & Emig

Carboniferous to Triassic

 China
 Russia
 Spain

A member of Lingulata belonging to the group Lingulida and the family Lingulidae. The type species is "Lingularia" siberica Biernat & Emig (1993).

Experilingula larga[64]

Sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A member of Obolidae.

Flexaria magna[63]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Globiella kamiyassensis[74]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan

Glossella cuyanica[68]

Sp. nov

Valid

Lavié & Benedetto

Ordovician (Darriwilian)

San Juan Formation

 Argentina

Gondwanorthis[75]

Gen. et comb. nov

Valid[76]

Benedetto & Muñoz

Early Ordovician

 Argentina
 Iran

A plectorthoid brachiopod. A new genus for "Nanorthis" calderensis Benedetto (2007); genus also includes "Nanorthis" bastamensis Ghobadi Pour, Kebriaee-Zadeh & Popov (2011).

Gowanella[77]

Gen. et sp. nov

Valid

Hiller

Late Cretaceous (probably Maastrichtian)

Broken River Formation

 New Zealand

A member of Terebratulida related to Ostreathyris. The type species is G. capralis.

Grandispirifer qaidamensis[78]

Sp. nov

Valid

Lee, Shi & Chen in Shi et al.

Carboniferous (Serpukhovian)

Huaitoulata Formation

 China

A member of Spiriferoidea belonging to the family Spiriferidae.

Gretchispirifer[69]

Gen. et sp. nov

Valid

Oleneva

Devonian (EmsianEifelian)

Sharur Formation

 Azerbaijan

A member of Spiriferida belonging to the family Adolfiidae. The type species is G. dagnensis.

Gundaria[79]

Gen. et sp. nov

Valid

Angiolini et al.

Permian

 Tajikistan

The type species is G. insolita.

Harperoides[80]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

 United States
( Alaska)

A member of Strophomenida belonging to the subfamily Mesodouvillininae. The type species is Harperoides alaskensis.

Hemileurus politus[79]

Sp. nov

Valid

Angiolini et al.

Permian

 Tajikistan

Holcothyris campbelli[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Hubeinomena[65]

Gen. et sp. nov

Disputed

Zeng,Chen & Han in Zeng et al.

Ordovician (Hirnantian)

 China

A member of Strophomenida belonging to the family Sinomenidae. Genus includes new species H. wangjiawanensis. Rong et al. (2019) considered it to be a junior synonym of Coolinia.[81]

Hustedia shumardi[71]

Sp. nov

Valid

Torres-Martínez, Sour-Tovar & Barragán

Permian (Leonardian)

Paso Hondo Formation

 Mexico

Inflatia coodzavuii[63]

Sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (late Mississippian, middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea.

Iridistrophia (Flabellistrophia)[82]

Subgen. et comb. sp. nov

Valid

Jansen

Devonian (late Emsian to Eifelian)

 Germany
 Venezuela?

A member of Chilidiopsidae; a subgenus of Iridistrophia. The type species is "Orthis" hipponyx Schnur (1851); the subgenus also includes new species Iridistrophia (Flabellistrophia) musculosa and possibly also "Orthis" undifera Schnur (1853) and Iridistrophia dendritica Benedetto (1984).

Ishimia inflata[59]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Leptellinidae.

Isogramma nakamurai[74]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan

Jakutoproductus lenensis[83]

Sp. nov

Valid

Makoshin

Early Permian

 Russia

Jakutoproductus talchanensis[83]

Sp. nov

Valid

Makoshin

Early Permian

 Russia

Kjaerina (Kjaerina) gondwanensis[84]

Sp. nov

Valid

Colmenar

Ordovician (late SandbianKatian)

Gabian Formation
Glauzy Formation
Louredo Formation
Portixeddu Formation

 France
 Italy
 Portugal

A rafinesquinid strophomenid brachiopod, a species of Kjaerina.

Kjaerina (Villasina)[84]

Subgen. et 3 sp. et comb. nov

Valid

Colmenar

Ordovician (Katian)

Cavá Formation
Gabian Formation
Portilla de Luna Limestones
Portixeddu Formation
Porto de Santa Anna Formation
Punta Serpeddi Formation
Rosan Formation

 France
 Italy
 Portugal
 Spain

A rafinesquinid strophomenid brachiopod, a subgenus of Kjaerina. The type species of the subgenus is Kjaerina (Villasina) pedronaensis; the subgenus also contains "Hedstroemina" almadenensis Villas (1995), as well as new species Kjaerina (Villasina) meloui and Kjaerina (Villasina) pyrenaica.

Koneviella? fuscina[64]

Sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A lingulate brachiopod belonging to the family Zhanatellidae.

Kutchithyris challinori[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Kutchithyris waitomoensis[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Lacunites jaroslavi[85]

Sp. nov

Valid

Mergl & Kraft

Early Ordovician

Klabava Formation

 Czech Republic

A paterinate brachiopod.

Lampazarorthis[75]

Gen. et comb. et sp. nov

Valid[76]

Benedetto & Muñoz

Early Ordovician

 Argentina

A plectorthoid brachiopod. A new genus for "Eoorthis" bifurcata Harrington (1937); genus also includes "Nanorthis" brachymyaria Benedetto in Benedetto & Carrasco (2002), as well as new species Lampazarorthis alata.

Lepidomena multiplicata[59]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Strophomenida belonging to the family Leptellinidae.

Leptathyris gornensis[60]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Athyrididae.

Liaous[86]

Gen. et sp. nov

Valid

He & Chen in He et al.

Middle Triassic (early Anisian)

Xinyuan Formation

 China

A relative of Mentzelia and Paramentzelia. The type species is Liaous shaiwensis.

Loboidothyris awakinoensis[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Loboidothyris grantmackiei[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Loboidothyris marokopaensis[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Lyonia rochacamposi[87]

Sp. nov

Valid

Taboada et al.

Early Permian (latest Asselian-earliest Sakmarian)

Taciba Formation

 Brazil

A member of Productida belonging to the family Auriculispinidae, a species of Lyonia.

Mendozotreta[88]

Gen. et comb. nov

Valid

Holmer et al.

Ordovician

Antelope Valley Limestone
Lindero Formation

 Argentina
 United States
( Nevada)

A member of Acrotretida belonging to the family Acrotretidae. The type species is "Conotreta" devota Krause & Rowell (1975).

Mesoleptostrophia belli[89]

Sp. nov

Valid

Earp

Early Devonian

Montys Hut Formation

 Australia

Minutella bulgarica[90]

Sp. nov

Valid

Bitner & Motchurova-Dekova

Miocene (Badenian)

 Bulgaria

Minutomena[65]

Gen. et sp. nov

Valid

Zeng,Zhang & Han in Zeng et al.

Ordovician (Hirnantian)

 China

A member of the family Strophomenidae. Genus includes new species M. yichangensis.

Minutorthis[65]

Gen. nov

Valid

Zeng,Chen & Zhang in Zeng et al.

Ordovician (Hirnantian)

 China

A member of Orthida belonging to the family Toxorthidae.

Monelasmina montisjosephi[60]

Sp. nov

Valid

Baliński in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Draboviidae.

Nasakia[91]

Gen. et sp. nov

Valid

Streng et al.

Cambrian

Henson Gletscher Formation

 Greenland

A member of Rhynchonelliformea belonging to the class Obolellata and the order Naukatida. The type species is Nasakia thulensis.

Numericoma rowelli[88]

Sp. nov

Valid

Holmer et al.

Ordovician (Darriwilian)

Antelope Valley Limestone
Ponon Trehue Formation

 Argentina
 United States
( Nevada)

A member of Lingulata belonging to the family Ephippelasmatidae.

Nushbiella kleithria[55]

Sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Siphonotretidae.

Obliquorhynchia[92]

Gen. et comb. nov

Valid

Schrøder, Lauridsen & Surlyk

Paleocene (Danian)

Faxe Formation
Vigny Formation

 Denmark
 France
 Sweden

A member of Rhynchonellida belonging to the superfamily Pugnacoidea and the family Basiliolidae; a new genus for "Terebratula" flustracea von Buch (1834).

Orthis dehmollaensis[59]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Orthida belonging to the family Orthidae.

Paramirorthis[65]

Gen. et sp. nov

Disputed

Zeng,Wang & Peng in Zeng et al.

Ordovician (Hirnantian)

 China

A member of the family Dalmanellidae. Genus includes new species P. minuta. Rong et al. (2019) considered it to be a junior synonym of Mirorthis.[81]

Paraspirifer (Laurentispirifer)[82]

Subgen. et comb. nov

Valid

Jansen

Middle Devonian

 United States
 Venezuela

A subgenus of Paraspirifer. The type species is Paraspirifer conradi Godefroid & Fagerstrom (1983); the subgenus also includes "Delthyris" acuminata Conrad (1839), "Terebratula" acuminatissima de Castelnau (1843), "Spirifer" bownockeri Stewart (1927), Paraspirifer halli Godefroid & Fagerstrom (1983) and Paraspirifer clarkei Godefroid & Fagerstrom (1983).

Paraspirifer (Mosellospirifer)[82]

Subgen. et comb. nov

Valid

Jansen

Devonian (late Emsian to early Eifelian

 Germany
 Canada?
 China?

A subgenus of Paraspirifer. The type species is Paraspirifer sandbergeri Solle (1971); the subgenus also includes Spirifer auriculatus Sandberger & Sandberger (1856), Paraspirifer sandbergeri longimargo Solle (1971) (elevated to species rank), Paraspirifer eos Solle (1971) and Paraspirifer sandbergeri nepos Solle (1971). The subgenus might also include Paraspirifer gigantea Su (1976) and Paraspirifer desbiensi Bizzarro & Lespérance (1999).

Parazhanatella[64]

Gen. et sp. nov

Valid

Percival, Engelbretsen & Peng

Cambrian

Huaqiao Formation

 China

A lingulate brachiopod belonging to the family Zhanatellidae. The type species is P. paibia.

Pedderia[80]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

 United States
( Alaska)

A member of Rhynchonellida belonging to the family Pygmaellidae. The type species is Pedderia fragosa.

Phragmorthis shahrudensis[59]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Orthida belonging to the family Phragmorthidae.

Prospira pseudostruniana[93]

Sp. nov

Valid

Mottequin & Brice

Devonian (late Famennian)

Etrœungt Formation

 France

A member of Spiriferida belonging to the family Spiriferidae.

Psygmakantha[55]

Gen. et sp. nov

Valid

Percival in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Ephippelasmatidae. The type species is P. malachiensis.

Punctospirifer gerankalasus[69]

Sp. nov

Valid

Oleneva

Carboniferous (Tournaisian)

 Azerbaijan

A member of Spiriferinida belonging to the family Punctospiriferidae.

Qaidamospirifer[78]

Gen. et sp. nov

Valid

Chen, Lee & Shi in Shi et al.

Carboniferous (Serpukhovian)

Huaitoulata Formation

 China

A member of Spiriferoidea belonging to the family Choristitidae. The type species is Q. elongatus.

Rafinesquina (Mesogeina)[84]

Subgen. et comb. et 2 sp. nov

Valid

Colmenar

Ordovician (Katian)

Bohdalec Formation
Fombuena Formation
Gabian Formation
Lower Ktaoua Formation
Porto de Santa Anna Formation
Upper Tiouririne Formation
Zahorany Formation

 Czech Republic
 France
 Germany
 Morocco
 Portugal
 Spain

A rafinesquinid strophomenid brachiopod, a subgenus of Rafinesquina. The type species of the subgenus is "Leptaena" pseudoloricata Barrande (1848); the subgenus also includes Rafinesquina pomoides Havlíček (1971), as well as new species Rafinesquina (Mesogeina) gabianensis and Rafinesquina (Mesogeina) loredensis.

Rhipidomella magna[74]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan

Rogorthis? oriens[59]

Sp. nov

Valid

Popov, Kebriaee-Zadeh & Pour

Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Orthida belonging to the family Orthidae.

Sampo suduvensis[70]

Sp. nov

Valid

Paškevičius & Hints

Late Ordovician

 Lithuania
 Russia
( Kaliningrad Oblast)

A member of Strophomenida belonging to the family Leptestiidae.

Sartenaerirhynchus[82]

Gen. et comb. nov

Valid

Jansen

Devonian (middle Siegenian to late Emsian)

 Belgium
 Germany
 Luxembourg

A member of Rhynchonellida belonging to the superfamily Uncinuloidea. The type species is "Terebratula" antiqua Schnur (1853); genus also includes "Uncinulus" frontecostatus Drevermann (1902)

Scaphelasma quadratum[55]

Sp. nov

Valid

Engelbretsen in Percival et al.

Ordovician

 Australia

A member of Lingulata belonging to the family Scaphelasmatidae.

Schizocrania equestra[94]

Sp. nov

Valid

Mergl & Nolčová

Ordovician (Katian)

Bohdalec Formation

 Czech Republic

A member of Discinoidea belonging to the family Trematidae.

Septaliphoria felberi[61]

Sp. nov

Valid

Sulser

Middle Jurassic (Callovian)

  Switzerland

A member of Rhynchonellida belonging to the superfamily Hemithiridoidea and the family Cyclothyrididae.

Siberioproductus[67]

Gen. et sp. nov

Valid

Baranov, Sokiran & Blodgett

Devonian (Famennian)

Chekursk Formation

 Russia
( Sakha Republic)

A member of Rhynchonelliformea belonging to the group Productida and the family Productellidae. The type species is S. boreus.

Sinomena[65]

Gen. et sp. nov

Disputed

Zeng,Chen & Han in Zeng et al.

Ordovician (Hirnantian)

 China

A member of Strophomenida belonging to the family Sinomenidae. Genus includes new species S. typica. Rong et al. (2019) considered it to be a junior synonym of Eostropheodonta.[81]

Skenidioides cretus[60]

Sp. nov

Valid

Halamski in Baliński, Racki & Halamski

Devonian (Frasnian)

 Poland

A member of Skenidiidae.

Spinobolus[95]

Gen. et sp. nov

Valid

Zhang et al.

Cambrian

Shuijingtuo Formation

 China

A linguloid brachiopod. The type species is Spinobolus popovi.

Tapongaspirifer[89]

Gen. et sp. nov

Valid

Earp

Early Devonian

Montys Hut Formation

 Australia

A member of Spiriferida belonging to the family Filispiriferidae. The type species is Tapongaspirifer melodiae.

Tasmanospirifer[57][58]

Gen. et sp. et comb. nov

Valid

Waterhouse

Permian

 Australia

A member of Spiriferida. Genus includes new species T. clarkei, as well as "Spirifer" tasmaniensis Morris (1845).

Taungurungetes[89]

Gen. et comb. nov

Valid

Earp

Early Devonian

Montys Hut Formation
Norton Gully Sandstone

 Australia

A possible member of Devonochonetinae. The type species is "Chonetes" taggertyensis Gill (1945) from the Montys Hut Formation; genus also contains second, unnamed species from the Norton Gully Sandstone.

Terebratulina leeae[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Terebratulina putiensis[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Thaerodonta notabile[96]

Sp. nov

Valid

Paškevičius

Ordovician (Katian)

Kaimynai Formation

 Lithuania

Thulatrypa[97]

Gen. et sp. et comb. nov

Valid

Huang et al.

Silurian

 China
 Norway

An atrypoid brachiopod. The type species is Thulatrypa gregaria; genus also contains "Meifodia" orientalis Rong, Xu & Yang (1974).

Tomteluva[91]

Gen. et sp. nov

Valid

Streng et al.

Cambrian

Stephen Formation

 Canada
( British Columbia)

A member of Rhynchonelliformea belonging to the class Obolellata and the order Naukatida. The type species is Tomteluva perturbata.

Trentingula[98]

Gen. et 3 sp. et comb. nov

Valid

Posenato

Late Permian and Early Triassic

 Hungary
 Italy
 United States
( Wyoming)

A member of Linguloidea belonging to the family Lingulidae. The type species is T. lorigae; genus also includes new species T. mazzinensis and T. prinothi, as well as Trentingula borealis (Bittner, 1899).

Triangulospirifer[78]

Nom. nov

Valid

Lee in Shi et al.

Carboniferous

 United Kingdom

A member of Spiriferida. A replacement name for Triangularia Poletaev (2001).

Trimena[65]

Gen. nov

Valid

Zeng,Wang & Peng in Zeng et al.

Ordovician (Hirnantian)

 China

A member of Plectambonitoidea belonging to the family Xenambonitidae.

Waddingtonia[57]

Gen. et comb. nov

Valid

Waterhouse

Devonian

Delorme Formation

 Canada
( Yukon)

A member of Spiriferida. The type species is "Reticulariopsis" warreni Perry (1984).

Weberproductus[63]

Gen. et sp. nov

Valid

Torres-Martínez & Sour-Tovar

Carboniferous (middle Pennsylvanian)

Ixtaltepec Formation

 Mexico

A member of Productoidea. Genus includes new species W. donajiae.

Wenjukovispirifer[69]

Gen. et comb. nov

Valid

Oleneva

Devonian (Famennian)

 Russia

A member of Spiriferida belonging to the family Cyrtospiriferidae. The type species is "Spirifer" koscharicus Ljaschenko (1959); genus also includes "Spirifer" archiaci Murchison (1840), "Spirifer" brodi Wenjukov (1886) and W. lebedjanicus (Nalivkin, 1947).

Yichangomena[65]

Gen. et sp. nov

Disputed

Zeng,Zhang & Han in Zeng et al.

Ordovician (Hirnantian)

 China

A member of Strophomenida belonging to the family Sinomenidae. Genus includes new species Y. dingjiapoensis. Rong et al. (2019) considered it to be a junior synonym of Eostropheodonta.[81]

Zeilleria opuatiaensis[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Zeilleria waiohipaensis[62]

Sp. nov

Valid

MacFarlan

Jurassic

 New Zealand

A member of Terebratulida.

Molluscs[edit]

Echinoderms[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Alternocidaris[99]

Gen. et comb. nov

Valid

El Qot, Abdelhamid, & Abdelghany

Late Cretaceous (Cenomanian, possibly Turonian)

 Egypt

A sea urchin belonging to the group Camarodonta and the family Glyphocyphidae. The type species is "Tiaridia" weldoni Fourtau (1921).

Applinocrinus russelli[100]

Sp. nov

Valid[101]

Gale

Late Cretaceous (Maastrichtian)

Prairie Bluff Formation

 United States
( Mississippi)

A crinoid belonging to the group Articulata and the family Saccocomidae.

Apsidocrinus doreckae[102]

Sp. nov

Valid

Konieczyński, Pisera & Fózy

Early Cretaceous

 Hungary

A cyrtocrinid crinoid, a species of Apsidocrinus.

Arabicodiadema[103]

Gen. et comb. et sp. nov

Valid

Abdelhamid, El Qot & Abdelghany

Cretaceous (Albian to Cenomanian)

 Oman
 United Arab Emirates

A heterodiadematid sea urchin. The type species is "Trochodiadema" dhofarense Roman (1991); genus also includes new species Arabicodiadema alii.

Atalopegaster[104]

Gen. et sp. nov

Valid

Blake & Guensburg

Late Jurassic (Oxfordian)

Swift Formation

 United States
( Montana)

A starfish belonging to the family Stichasteridae. The type species is A. gundersoni.

Bystrowicrinus (col.) depressus[105]

Sp. nov

Valid

Donovan & Keighley

Late Silurian

West Point Formation

 Canada
( Quebec)

A crinoid.

Calliderma lindneri[106]

Sp. nov

Valid

Niebuhr & Seibertz

Late Cretaceous (Turonian)

Schmilka Formation

 Germany

A starfish belonging to the family Goniasteridae, a species of Calliderma.

Carolinaster[107]

Gen. et sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Castle Hayne Limestone

 United States
( North Carolina)

A heart urchin belonging to the family Prenasteridae. Genus includes new species C. varnami.

Comptoniaster michaelisi[106][108]

Nom. nov

Valid

Niebuhr & Seibertz

Late Cretaceous (middle Turonian to middle Coniacian)

 Czech Republic
 Germany
 Poland

A starfish belonging to the family Goniasteridae; a replacement name for Asterias schulzii Cotta sensu Roemer (1840, 1841).

Costatocrinus[100]

Gen. et 2 sp. nov

Valid[101]

Gale

Late Cretaceous (Campanian)

Taylor Formation

 United Kingdom
 United States
( Texas)

A crinoid belonging to the group Articulata and the family Saccocomidae. The type species is C. brydonei; genus also includes C. mortimorei.

Cultellacrinus[100]

Gen. et 2 sp. nov

Valid[101]

Gale

Late Cretaceous

 Sweden
 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is C. gladius; genus also includes C. labyrinthus.

Cunidentechinus[109]

Gen. et comb. nov

Valid

Smith

Early Jurassic

 United Kingdom

A stem-carinacean sea urchin; a new genus for "Echinus" minutus Buckman in Murchison (1845).

Darwinaster[110]

Gen. et sp. nov

Valid

Hunter, Rushton & Stone

Early Devonian

Fox Bay Formation

 Falkland Islands

A brittle star related to members of the genus Protaster. Genus includes new species D. coleenbiggsae.

Echinolampas andalusiensis[107]

Sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Moodys Branch Formation

 United States
( Alabama)

A sea urchin belonging to the family Echinolampadidae.

Gitolampas twitchelli[107]

Sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Santee Limestone

 United States
( South Carolina)

A sea urchin belonging to the group Cassiduloida.

Glyptocrinus nodosus[111]

Sp. nov

Valid

Kallmeyer & Ausich

Ordovician (Katian)

Kope Formation

 United States
( Kentucky
 Ohio)

A camerate crinoid, a species of Glyptocrinus.

Goniopygus macrotuberculatus[99]

Sp. nov

Valid

El Qot, Abdelhamid, & Abdelghany

Late Cretaceous (Cenomanian)

Raha Formation

 Egypt

A sea urchin belonging to the group Arbacioida and the family Acropeltidae.

Goniopygus subaequalis[99]

Sp. nov

Valid

El Qot, Abdelhamid, & Abdelghany

Late Cretaceous (Cenomanian)

Raha Formation

 Egypt

A sea urchin belonging to the group Arbacioida and the family Acropeltidae.

Haimea brooksi[107]

Sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Ocala Limestone

 United States
( Florida)

A sea urchin belonging to the group Oligopygoida.

Hessicrinus[100]

Gen. et 2 sp. nov

Valid[101]

Gale

Late Cretaceous (Campanian)

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is H. filigree; genus also includes H. scalaensis.

Hiiumaacrinus[112]

Gen. et sp. nov

Valid

Ausich & Wilson

Silurian (Rhuddanian)

Hilliste Formation

 Estonia

A crinoid. Genus includes new species H. vinni.

Jakeocrinus[100]

Gen. et sp. nov

Valid[101]

Gale

Late Cretaceous (early Campanian)

Taylor Formation

 United States
( Texas)

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is J. ellisensis.

Lakotacrinus[113]

Gen. et sp. nov

Valid

Hunter et al.

Late Cretaceous (Campanian)

Pierre Shale

 United States
( South Dakota)

A stalked crinoid. The type species is L. brezinai.

Lehmannaster[114]

Gen. et sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (late Darriwilian)

Traveusot Formation

 France

A stenuroid asterozoan. Genus includes new species L. spinosus.

Lopidiaster[114]

Gen. et sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (Floian)

Garden City Formation

 United States
( Utah)

A stenuroid asterozoan. Genus includes new species L. jamisoni.

Loriolidiadema[103]

Gen. et comb. nov

Valid

Abdelhamid, El Qot & Abdelghany

Cretaceous (Aptian-Cenomanian)

 Egypt
 Portugal
Border region between  Syria and  Lebanon

A heterodiadematid sea urchin. The type species is "Pseudodiadema" libanoticum de Loriol (1887); genus also includes Loriolidiadema sculptile (de Loriol, 1887).

Lucernacrinus[100]

Gen. et sp. nov

Valid[101]

Gale

Late Cretaceous

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is L. woodi.

Maennilocystis[115]

Gen. et sp. nov

Valid

Paul & Rozhnov

Late Ordovician

 Estonia

A cystoid belonging to the family Callocystitidae. The type species is M. heckeri.

Manfredaster praebulbiferus[106]

Sp. nov

Valid

Niebuhr & Seibertz

Late Cretaceous (Cenomanian to Coniacian)

Dölzschen Formation
Schrammstein Formation
Strehlen Formation

 Germany

A starfish belonging to the group Valvatida and the family Stauranderasteridae.

Mariania comaschicariae[116]

Sp. nov

Valid

Stara, Borghi & Kroh

Miocene (Aquitanian to early Burdigalian)

Nurallao Formation

 Italy

A heart urchin belonging to the superfamily Spatangoidea, a species of Mariania.

Mariania stefaninii[116]

Sp. nov

Valid

Stara, Borghi & Kroh

Miocene (late Burdigalian to early Langhian)

Pantano Formation

 Italy

A heart urchin belonging to the superfamily Spatangoidea, a species of Mariania.

Meturaster[114]

Gen. et sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (middle Darriwilian)

Lehman Formation

 United States
( Nevada)

A starfish. Genus includes new species M. belli.

Monophoraster telfordi[117]

Sp. nov

Valid

Mooi, Martínez & del Río

Early Miocene

Chenque Formation

 Argentina

A sand dollar.

Nucleolites usoi[118]

Sp. nov

Valid

Forner i Valls

Early Cretaceous (Barremian)

Artoles Formation

 Spain

A sea urchin.

Osteocrinus sinensis[119]

Sp. nov

Valid

Hess, Etter & Hagdorn

Late Triassic (early Carnian)

Xiaowa Formation

 China

A crinoid belonging to the group Roveacrinida, a species of Osteocrinus.

Pentamerocrinus kosovi[120]

Sp. nov

Valid

Rozhnov

Ordovician

 Russia
( Leningrad Oblast)

A crinoid belonging to the group Inadunata, a species of Pentamerocrinus.

Petraster crozonensis[114]

Sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (early Katian)

Kermeur Formation

 France

A starfish.

Phyllocrinus bersekensis[102]

Sp. nov

Valid

Konieczyński, Pisera & Fózy

Early Cretaceous

 Hungary

A cyrtocrinid crinoid, a species of Phyllocrinus.

Placentinechinus[121]

Gen. et sp. nov

Valid[122]

Borghi & Garilli

Pleistocene (GelasianCalabrian)

 Italy

A sea urchin belonging to the family Temnopleuridae. The type species is P. davolii.

Plagiochasma saurai[118]

Sp. nov

Valid

Forner i Valls

Early Cretaceous (Barremian)

Artoles Formation

 Spain

A sea urchin.

Platelicrinus longispinus[100]

Sp. nov

Valid[101]

Gale

Late Cretaceous

 United Kingdom

A crinoid belonging to the group Articulata and the family Roveacrinidae.

Protaeropsis[123]

Nom. nov

Valid

Doweld

Paleocene (Thanetian)

 Spain

A heart urchin; a replacement name for Sphenaster Jeffery in Smith et al. (1999).

Protaxocrinus estoniensis[112]

Sp. nov

Valid

Ausich & Wilson

Silurian (Rhuddanian)

Hilliste Formation

 Estonia

A crinoid.

Pseudomilnia[109]

Gen. et comb. nov

Valid

Smith

Middle Jurassic

 France
 United Kingdom

A salenioidan sea urchin; a new genus for "Acrosalenia" miliaria Paris (1908).

Sagittacrinus[100]

Gen. et sp. nov

Valid[101]

Gale

Late Cretaceous (early Campanian)

Taylor Formation

 United Kingdom
 United States
( Texas)

A crinoid belonging to the group Articulata and the family Saccocomidae. The type species is S. torpedo.

Salenia buddyi[107]

Sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene

Santee Limestone

 United States
( South Carolina)

A sea urchin belonging to the group Salenioida.

Schuchertia sprinklei[114]

Sp. nov

Valid

Blake, Guensburg & Lefebvre

Ordovician (Dapingian)

Kanosh Shale

 United States
( Utah)

A starfish.

Stellacrinus[100]

Gen. et 2 sp. nov

Valid[101]

Gale

Late Cretaceous (Santonian and Campanian)

Gulpen Formation
Taylor Formation
Vaals Formation

 Netherlands
 Sweden
 United Kingdom
 United States
( Texas)

A crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is S. hugesae; genus also includes S. pannosus.

Stomechinus phillipsii[109]

Sp. nov

Valid

Smith

Jurassic

 United Kingdom

A sea urchin.

Thinocrinus akanthos[124]

Sp. nov

Valid

Rhenberg, Ausich & Meyer

Carboniferous (Mississippian)

Fort Payne Formation

 United States
( Kentucky)

A camerate crinoid belonging to the group Monobathrida and the family Actinocrinitidae.

Trochotiara aalensis[109]

Sp. nov

Valid

Smith

Middle Jurassic

 United Kingdom

A sea urchin.

Wrightechnus[109]

Gen. et comb. nov

Valid

Smith

Middle Jurassic

 France
 United Kingdom

A sea urchin, possibly a stem-salenioidan; a new genus for "Acrosalenia" lycetti Wright (1851).

Conodonts[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Amydrotaxis corniculans truncus[125]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Cypricriodus[126][127]

Gen. et comb. nov

Valid

Murphy, Carls & Valenzuela-Ríos

Silurian and Early Devonian

 Australia
 United States
( Nevada)

Genus includes "Caudicriodus" hesperius (Klapper & Murphy, 1975).

Eognathodus kuangi[128]

Sp. nov

Valid

Earp

Early Devonian

 China

Eognathodus sulcatus lanei[125]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Eognathodus sulcatus sicatus[125]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Booth Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Gladigondolella okayi[129]

Sp. nov

Valid

Kiliç

Triassic

Kayabaşı Limestone

 Turkey

Gondolella wardlawi[130]

Sp. nov

Valid

Nestell & Pope in Nestell, Wardlaw & Pope

Carboniferous (Pennsylvanian)

Excello Shale

 United States
( Iowa)

Heliagnathus[125]

Gen. et sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida of uncertain phylogenetic placement. The type species is H. parvilabiatus.

Icriodus ampliatus[125]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Prioniodontida belonging to the family Icriodontidae.

Icriodus michiganus[131]

Sp. nov

Valid

Narkiewicz & Bultynck

Devonian (Eifelian)

Michigan Basin

 United States

Icriodus praealternatus ferus[132]

Subsp. nov

Valid

Wang et al.

Late Devonian (Frasnian/Famennian boundary)

Hongguleleng Formation

 China

A subspecies of Icriodus praealternatus.

Icriodus plurinodosus[132]

Sp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

A species of Icriodus.

Icriodus stenoancylus junggarensis[132]

Subsp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

A subspecies of Icriodus stenoancylus.

Idiognathodus lateralis[133]

Sp. nov

Valid

Hogancamp, Barrick & Strauss

Carboniferous (early Gzhelian)

 China
 Russia
 Ukraine
 United States

Originally described as a species of Idiognathodus, but subsequently transferred to the genus Heckelina.[134]

Idiognathodus tuberis[130]

Sp. nov

Valid

Nestell, Wardlaw & Pope

Carboniferous (Pennsylvanian)

Excello Shale

 United States
( Iowa)

A member of Ozarkodinida belonging to the family Idiognathodontidae.

Neogondolella cuneiforme[135]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)

Neogondolella curva[135]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella dilacerata[135]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)

Originally described as a species of Neogondolella, but subsequently transferred to the genus Magnigondolella.[136]

Neogondolella hastata[135]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella indicta[135]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella panlaurentia[135]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella tenera[135]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neogondolella vellicata[135]

Sp. nov

Valid

Golding & Orchard

Middle Triassic (Anisian)

 Canada
( British Columbia)
 United States
( Nevada)

Neopolygnathus huijunae[132]

Sp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

A species of Neopolygnathus.

Neospathodus planus[137]

Sp. nov

Valid

Chen et al.

Early Triassic

 Slovenia

Originally described as a species of Neospathodus; Bai et al. (2021) argued that it should be reassigned to the genus Triassospathodus.[138]

Oulodus astriatus[125]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Booth Limestone

 Australia

A member of Prioniodinida belonging to the family Prioniodinidae.

Ozarkodina multistriola[125]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina scoliciformis[125]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina selfi cordata[125]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina selfi lanceola[125]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Ozarkodina selfi lenticula[125]

Subsp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of Ozarkodinida belonging to the family Spathognathodontidae.

Panderodus rhytiodus[125]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Mountain Dam Limestone

 Australia

A member of the family Panderodontidae.

Pelekysgnathus inequalis[125]

Sp. nov

Valid

Mathieson et al.

Early Devonian

Booth Limestone

 Australia

A member of Prioniodontida belonging to the family Icriodontidae.

Pelekysgnathus jeppssoni[139]

Sp. nov

Valid

Nazarova

Devonian (Eifelian)

 Russia
( Bryansk Oblast)

Platyvillosus corniger[137]

Sp. nov

Valid

Chen et al.

Early Triassic

 Slovenia

Polygnathus abaimovae[140]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus beckeri[140]

Nom. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

A replacement name for Polygnathus inflexus Baranov (1992).

Polygnathus datnensis[140]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus dogdensis[140]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

Polygnathus novozemelicus[140]

Sp. nov

Valid

Baranov & Blodgett

Devonian (Emsian)

 Russia

"Polygnathus" pseudocommunis[132]

Sp. nov

Valid

Wang et al.

Late Devonian (early Famennian)

Hongguleleng Formation

 China

Possibly a species of Polygnathus.

Polygnathus s

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