2021 in paleomalacology

List of years in paleomalacology
In science
2018
2019
2020
2021
2022
2023
2024
In paleontology
2018
2019
2020
2021
2022
2023
2024
In paleomammalogy
2018
2019
2020
2021
2022
2023
2024
In paleobotany
2018
2019
2020
2021
2022
2023
2024
In arthropod paleontology
2018
2019
2020
2021
2022
2023
2024
In paleoentomology
2018
2019
2020
2021
2022
2023
2024
In paleoichthyology
2018
2019
2020
2021
2022
2023
2024
In reptile paleontology
2018
2019
2020
2021
2022
2023
2024
In archosaur paleontology
2018
2019
2020
2021
2022
2023
2024

This list of fossil molluscs described in 2021 is a list of new taxa of fossil molluscs that were described during the year 2021, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2021.

Ammonites[edit]

New taxa[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Abichites infirmis[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Abichites ovalis[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Agathiceras iranicum[2]

Sp. nov

Valid

Korn & Ghaderi in Korn et al.

Carboniferous-Permian boundary

 Iran

Alienisphinctes[3]

Gen. et sp. nov

Valid

Énay & Mangold

Late Jurassic (Kimmeridgian)

Jubaila Formation

 Saudi Arabia

A member of the family Ataxioceratidae. The type species is A. cavelieri.

Amaltheus orientalis[4]

Sp. nov

Valid

Nakada et al.

Early Jurassic (Pliensbachian)

Kuruma Group

 Japan

Arasella falcata[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

An ammonite of uncertain placement, probably a member of the family Xenodiscidae.

Arcestes lawsi[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

Arnioceras ritterbushi[6]

Sp. nov

Valid

Taylor & Guex

Early Jurassic (Sinemurian)

Sunrise Formation

 United States
( Nevada)

Arnioceras sparsum[6]

Sp. nov

Valid

Taylor & Guex

Early Jurassic (Sinemurian)

Sunrise Formation

 United States
( Nevada)

Artyomites[7]

Gen. et sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Palaeophyllitidae. Genus includes new species A. antiquus.

Aulatornoceras frenklerae[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Aulatornoceras steinhauseni[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Aulatornoceras ventrosulcatum[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Boucaulticeras hawthornensis[6]

Sp. nov

Valid

Taylor & Guex

Early Jurassic (Sinemurian)

Sunrise Formation

 United States
( Nevada)

A member of the family Schlotheimiidae.

Bullatimorphites szontaghi[9]

Sp. nov

Valid

Galácz

Caloceras avus[10]

Sp. nov

Valid

Kment

Early Jurassic (Hettangian)

Catrisimoceras[11]

Gen. et comb. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and possibly to the subfamily Lytogyroceratinae. The type species is "Simoceras" catrianum Zittel (1870).

Cheltonia howarthi[12]

Sp. nov

Valid

Simms & Edmunds

Early Jurassic

 United Kingdom

Cibolaites petraschecki[13]

Sp. nov

Valid

Wilmsen et al.

Late Cretaceous (Turonian)

Brießnitz Formation

 Germany

Clivotirolites[1]

Gen. et 2 sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Dzhulfitidae. The type species is C. decoratus; genus also includes C. petilus.

Crassotornoceras hetzeneggeri[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Crassotornoceras nudum[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Crioceratites claveli[14]

Sp. nov

In press

Bert et al.

Early Cretaceous (Hauterivian)

Vocontian Basin

 France

Didymoceras morozumii[15]

Sp. nov

Valid

Misaki & Tsujino

Late Cretaceous (Campanian)

Hiketa Formation

 Japan

Dimorphodiscus[16]

Gen. et sp. et comb. nov

Valid

Vermeulen et al.

Early Cretaceous (Barremian)

 Algeria
Crimea
 France

A member of the family Holcodiscidae. The type species is D. lerisae; genus also includes D. uhligi (Karakash, 1907), D. sayni (Joleaud, 1912) and "Metahoplites" postnodosum Vermeulen et al. (2017).

Discoscaphites mullinaxorum[17]

Sp. nov

Valid

Witts et al.

Late Cretaceous (Maastrichtian)

Corsicana Formation

 United States
( Texas)

Domanikoceras martenbergense[18]

Sp. nov

Valid

Korn

Devonian (Frasnian)

 Germany

A member of the family Tornoceratidae.

Dzhulfites brevisellatus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Eoasianites baghevangensis[2]

Sp. nov

Valid

Korn & Ghaderi in Korn et al.

Carboniferous-Permian boundary

 Iran

Eoermoceras[3]

Gen. et 2 sp. nov

Valid

Énay & Mangold

Middle Jurassic (Bajocian)

Dhruma Formation

 Saudi Arabia

A member of the family Stephanoceratidae. The type species is E. glabrum; genus also includes new species E. angulatidomus.

Eolytoceras rohrbachensis[10]

Sp. nov

Valid

Kment

Early Jurassic (Hettangian)

Esfahanites[1]

Gen. et sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Dzhulfitidae. The type species is E. armatus.

Eutomoceras paladeji[19]

Sp. nov

Valid

Tongtherm, Nabhitabhata & Tuanapaya

Triassic

Chaiburi Formation

 Thailand

A member of the family Ceratitidae.

Eutomoceras wantanae[19]

Sp. nov

Valid

Tongtherm, Nabhitabhata & Tuanapaya

Triassic

Chaiburi Formation

 Thailand

A member of the family Ceratitidae.

Evopharciceras[20]

Gen. et sp. et comb. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany
 Morocco

A member of Agoniatitida belonging to the family Pharciceratidae. The type species is E. formosum; genus also includes "Pharciceras" evolvens Bockwinkel, Becker & Ebbighausen (2009).

Extropharciceras metallicum[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Galfettites orientalis[7]

Sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Galfettitidae.

Glyptophiceras mirum[7]

Sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Xenoceltitidae.

Gonioclymenia junggarensis[21]

Sp. nov

In press

Zong, Becker & Ma

Devonian (Famennian)

Hongguleleng Formation

 China

Hadrothisbites hanwangensis[22]

Sp. nov

In press

Mietto et al.

Late Triassic (Carnian)

Ma'antang Formation

 China

Holcodiscus pseudocaillaudi[16]

Sp. nov

Valid

Vermeulen et al.

Early Cretaceous (Barremian)

 France

A member of the family Holcodiscidae.

Hulenites kitamurai[23]

Sp. nov

In press

Matsukawa

Early Cretaceous (Aptian)

Todai Formation

 Japan

Inaptychoceras[23]

Gen. et sp. nov

In press

Matsukawa

Early Cretaceous (Aptian)

Todai Formation

 Japan

Genus includes new species I. koikei.

Kislovodskoceras[24]

Gen. et sp. nov

Valid

Polkovoy

Early Cretaceous (Aptian)

 Russia

A member of the family Desmoceratidae. The type species is K. kislovodskense.

Kurnubiella fallax[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Callovian)

Upper Dhruma Formation

 Saudi Arabia

A member of the family Pachyceratidae.

Laevisimoceras[11]

Gen. et sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is L. inornatum.

Linguatornoceras asbeckense[18]

Sp. nov

Valid

Korn

Devonian (Frasnian)

 Germany

A member of the family Tornoceratidae.

Lunupharciceras kochi[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Lutites[1]

Gen. et 5 sp. et comb. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Dzhulfitidae. The type species is L. lyriformis; genus also includes new species L. alius, L. paucis, L. plicatus and L. profundus, as well as Alibashites stepanovi Korn & Ghaderi in Korn et al. (2015).

Lytogyroceras tuberculatum[11]

Sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Lytogyroceratinae.

Maenioceras ornatum[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Maenioceratidae.

Nejdia duartei[25]

Sp. nov

Valid

Goy & Comas-Rengifo

Early Jurassic (Toarcian)

A member of the family Hildoceratidae.

Obtusicostites gigas[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Callovian)

Upper Dhruma Formation

 Saudi Arabia

A member of the family Perisphinctidae.

Ophimullericeras[26]

Gen. et sp. nov

Valid

Jenks et al.

Early Triassic

Dinwoody Formation

 United States
( Nevada)

A member of Ceratitida belonging to the group Meekocerataceae and the family Mullericeratidae. The type species is O. paullae.

Pachyerymnoceras arabicum[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Callovian)

Tuwaiq Mountain Limestone Formation

 Saudi Arabia

A member of the family Pachyceratidae.

Pachyerymnoceras magnum[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Callovian)

Tuwaiq Mountain Limestone Formation

 Saudi Arabia

A member of the family Pachyceratidae.

Palaeophyllites mishkini[7]

Sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Palaeophyllitidae.

Paracadoceras vasily[27]

Sp. nov

Valid

Gulyaev in Gulyaev & Ippolitov

Middle Jurassic (Callovian)

 Ukraine

Paracochloceras nunminensis[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

Parahoplites inaensis[23]

Sp. nov

In press

Matsukawa

Early Cretaceous (Aptian)

Todai Formation

 Japan

Parasimoceras[11]

Gen. et 2 sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is P. venturii; genus also includes P. pseudostenonis.

Paratirolites aduncus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Paratirolites baghukensis[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Paratirolites lanceolobatus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Paratirolites robustus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Paratirolites rubens[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Peripleurites gabbensis[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

A member of Choristocerataceae belonging to the family Rhabdoceratidae.

Pharciceras beyrichi[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Pharciceras ferrum[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Pharciceras kruegeri[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Placites heggi[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

A member of the family Gymnitidae.

Planisimoceras[11]

Gen. et sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is P. costotuberculatum.

Pluripharciceras ahlburgi[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Pronjaites yolkinensis[28]

Sp. nov

Valid

Mitta

Early Cretaceous

 Russia

Pseudogalfettites[7]

Gen. et sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Galfettitidae. Genus includes new species P. sakhnoi.

Psiloceras pseudocostosum[10]

Sp. nov

Valid

Kment

Early Jurassic (Hettangian)

Retrotornoceras juxi[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Rhacophyllites mulleri[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

Rhacophyllites volcanoensis[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

Rollierites biplicatum[29]

Sp. nov

Valid

Jain & Mazur

Middle Jurassic (Callovian)

 Poland

A member of the family Pachyceratidae.

Sandbergeroceras archiaci[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Triainoceratidae.

Saudicostites[3]

Gen. et sp. nov

Valid

Énay & Mangold

Middle Jurassic (Bathonian)

Dhruma Formation

 Saudi Arabia

A member of Perisphinctoidea of uncertain phylogenetic placement. The type species is S. arabicus.

Shevyrevites corrugatus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Xenodiscidae.

Sigaloceras fundator[27]

Sp. nov

Valid

Gulyaev in Gulyaev & Ippolitov

Middle Jurassic (Callovian)

 Ukraine

Sigaloceras kiselevi[27]

Sp. nov

Valid

Gulyaev in Gulyaev & Ippolitov

Middle Jurassic (Callovian)

 Ukraine

Simocosticeras[11]

Gen. et sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Simoceratinae. The type species is S. biornatum.

Sinotropites[22]

Gen. et sp. nov

In press

Mietto et al.

Late Triassic (Carnian)

Ma'antang Formation

 China

Genus includes new species S. sichuanensis.

Sonninia (Euhoploceras) crassicostae[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Bajocian)

Lower Dhruma Formation

 Saudi Arabia

A member of the family Sonniniidae.

Sonninia (Euhoploceras) turbator[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Bajocian)

Lower Dhruma Formation

 Saudi Arabia

A member of the family Sonniniidae.

Sonoraceras[30]

Gen. et sp. nov

In press

Samaniego-Pesqueira, Moreno-Bedmar & Álvarez-Sánchez

Early Cretaceous (Aptian)

 Mexico

Genus includes new species S. tepachensis.

Stenopharciceras lotzi[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Stoliczkaia (Lamnayella) duboisi[31]

Sp. nov

Valid

Kennedy & Latil

Late Cretaceous (Cenomanian)

 France

A member of the family Lyelliceratidae.

Stoyanowites parallelus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Dzhulfitidae.

Strenoceras djangurense[32]

Sp. nov

Valid

Mitta

Middle Jurassic (Bajocian)

 Russia

Sulcicostites[11]

Gen. et sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Simoceratinae. The type species is S. costatus.

Timorites dendroides[33]

Sp. nov

Valid

Korn & Ghaderi in Tabrizi et al.

Permian (Wuchiapingian)

Julfa Formation

 Iran

A member of the family Cyclolobidae.

Tornoceras aequilobum[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Tornoceras fundibulum[18]

Sp. nov

Valid

Korn

Devonian (Famennian)

A member of the family Tornoceratidae.

Tornoceras occasus[34]

Sp. nov

Valid

Korn

Late Devonian

 Morocco

A member of the family Tornoceratidae.

Tornoceras ortus[34]

Sp. nov

Valid

Korn

Late Devonian

 Morocco

A member of the family Tornoceratidae.

Tornoceras vretterense[18]

Sp. nov

Valid

Korn

Devonian (Givetian)

A member of the family Tornoceratidae.

Tropaeum evolutum[35]

Sp. nov

In press

Lehmann & Bayliss

Early Cretaceous (Aptian)

Hythe Formation

 United Kingdom

Tropidoceras komarovi[36]

Sp. nov

Valid

Zaitsev

Early Jurassic

Crimean Peninsula

Vicininodiceras dalriatense[12]

Sp. nov

Valid

Simms & Edmunds

Early Jurassic

 United Kingdom

Volanoceras cuneospinatum[11]

Sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Volanoceratinae.

Waehneroceras morawetzi[10]

Sp. nov

Valid

Kment

Early Jurassic (Hettangian)

Wordieoceras mullenae[26]

Sp. nov

Valid

Jenks et al.

Early Triassic

Dinwoody Formation

 United States
( Nevada)

A member of Ceratitida belonging to the group Meekocerataceae and the family Ophiceratidae.

Research[edit]

  • A modern review of the palaeobiology of heteromorph ammonoids is published by Hoffmann et al. (2021), including details of their anatomy, buoyancy, locomotion, predators, diet, palaeoecology, and extinction.[37]
  • A study on the impact of the Permian–Triassic extinction event on the morphological diversity of ammonites is published by Dai, Korn & Song (2021).[38]
  • A study on the morphology and ontogeny of whorl profiles of the late Anisian ceratitid ammonites is published by Bischof et al. (2021).[39]
  • An exceptionally preserved specimen of Sigaloceras enodatum, providing information on the anatomy of ammonite muscles, is described from the Middle Jurassic (Callovian) Kellaways Sand Member of Colne Gravel (Gloucestershire, United Kingdom) by Cherns et al. (2021), who evaluate the implications of this specimen for the knowledge of the swimming mechanism of ammonites.[40]
  • Soft parts of a perisphinctid belonging to the genus Subplanites, separated from the conch either taphonomically or during a failed predation, are described from the Tithonian conservation deposits of Wintershof (southern Germany) by Klug et al. (2021).[41]
  • The first physical compression experiments on model ammonite septa are performed by Johnson et al. (2021), who attempt to determine whether the increasingly complex fractal margins of ammonite septa might have evolved to increase resistance to shell-crushing predators.[42]
  • A study aiming to determine the function of complex septa in the ammonite shell, providing evidence of a relationship between septal complexity and capacity for liquid retention within chambers of the shell, likely improving buoyancy regulation, is published by Peterman et al. (2021).[43]
  • Two specimens of Peltoceras athleta having both female and male features are described from Callovian limestone beds in Méron near Montreuil-Bellay (France) by Frau & Boursicot (2021), who interpret these specimens as cases of intersexuality in ammonites.[44]
  • Soft tissue remains preserved within the conch of a specimen of Proleymeriella schrammeni from the Albian of Germany are described by Hoffmann et al. (2021).[45]
  • A study on the arrangement in space and probable nature of hook-like structures found with specimens of Rhaeboceras halli is published by Smith et al. (2021), who interpret these structures as representing part of the brachial crown armature, and propose a hypothetical reconstruction of an ammonite brachial crown.[46]
  • A study on the systematics and phylogeny of the genus Aegocrioceras is published by Weinkauf, Hoffmann & Wiedenroth (2021).[47]
  • A study on the hydrostatic properties of Baculites compressus, and on their implications for the knowledge of the swimming capabilities and ecology of this ammonite and orthoconic cephalopods in general, is published by Peterman & Ritterbush (2021).[48]
  • Revision of the fossil material of Parapuzosia seppenradensis, including historic specimens from Europe and the United States and new material from England and Mexico, and a study on the ontogeny, distribution and evolution of this ammonite, is published by Ifrim et al. (2021).[49]
  • A study on the habitat depth preferences of Hoploscaphites constrictus from the European Boreal Chalk Sea, as indicated by stable isotope thermometry of aptychi and by data from predation marks on the scaphitid moulds, is published by Machalski et al. (2021).[50]
  • A study on the relationship between sutural complexity and body size in Cretaceous ammonites is published by Pérez-Claros (2021).[51]

Other cephalopods[edit]

New taxa[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Acrocoelites ippolitae[52]

Sp. nov

Valid

Weis in Weis et al.

Middle Jurassic (Aalenian)

 Germany

A belemnite.

Anglonautilus hewaidyi[53]

Sp. nov

In press

Moneer, Bazeen & Kassab

Late Cretaceous (Maastrichtian)

 Egypt

A nautiloid cephalopod.

Angulithes monqarensis[53]

Sp. nov

In press

Moneer, Bazeen & Kassab

Late Cretaceous (Maastrichtian)

 Egypt

A nautiloid cephalopod.

Arionoceras tokushimaense[54]

Sp. nov

Valid

Niko

Silurian (Ludlow)

Suberidani Group

 Japan

Bactroceras fluvii[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Dapingian)

Valhallfonna Formation

 Norway

Boreopeltis ifrimae[56]

Sp. nov

In press

Fuchs & Stinnesbeck

Late Cretaceous (Turonian)

 Mexico

Buttsoceras buldrebreenense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Troedssonellidae.

Cycloplectoceras hinlopense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Tarphyceratidae.

Cyclostomiceras profilstrandense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Cyclostomiceratidae.

Deltoceras beluga[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Cyclostomiceratidae.

Eosomichelinoceras borealis[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Baltoceratidae.

Eosomichelinoceras submedius[57]

Sp. nov

Valid

Evans et al.

Ordovician (Darriwilian)

Pelmis Formation

 Iran

A member of the family Baltoceratidae.

Ethanoceras[55]

Gen. et sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A relative of Rioceras. The type species is E. solitudines.

Gzheloceras aisenvergi[58]

Sp. nov

Valid

Dernov

Carboniferous (Serpukhovian)

Samara Formation

 Ukraine

A member of the family Tainoceratidae.

Hemichoanella occulta[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Hinlopoceras[55]

Gen. et 2 sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A relative of Rioceras. The type species is H. tempestatis; genus also includes H. venti.

Jovellania praetermissa[59]

Sp. nov

Valid

Brauckmann & Koch

Devonian (Givetian)

Massenkalk Formation

 Germany

Knightoceras extorris[58]

Sp. nov

Valid

Dernov

Carboniferous (Pennsylvanian)

 Ukraine

A member of the family Koninckioceratidae.

Lawrenceoceras ebenus[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Lawrenceoceras larus[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Litoceras profilbekkenense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Mojsvaroceras gianii[60]

Sp. nov

Valid

Pieroni & Prinoth

Middle Triassic (Anisian)

 Italy

A nautiloid.

Nyfrieslandoceras[55]

Gen. et sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of Oncocerida belonging to the family Phthanoncoceratidae. The type species is N. bassleroceroides.

Olenidslettoceras[55]

Gen. et sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of Oncocerida belonging to the family Phthanoncoceratidae. The type species is O. farmi.

Paracenoceras waageni[61]

Sp. nov

Valid

Halder

Middle Jurassic (Callovian)

Chari Formation

 India

A member of the family Paracenoceratidae.

Planetoceras yefimenkoi[58]

Sp. nov

Valid

Dernov

Carboniferous (Bashkirian)

Mospinka Formation

 Ukraine

A member of the family Koninckioceratidae.

Proterocameroceras valhallfonnense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Protocycloceras minor[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Troedssonellidae.

Rhynchorthoceras yizanense[62]

Sp. nov

Valid

Fang in Fang et al.

Ordovician

 China

A member of Lituitida.

Sinoceras complexum[62]

Sp. nov

Valid

Fang in Fang et al.

Ordovician

 China

A member of Lituitida.

Sorosoceras[57]

Gen. et sp. nov

Valid

Evans et al.

Ordovician (Darriwilian)

Pelmis Formation

 Iran

A member of the family Geisonoceratidae. The type species is S. castellum.

Svalbardoceras[55]

Gen. et 2 sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A relative of Rioceras. The type species is S. sterna; genus also includes S. skua.

Tyrioceras longicameratum[62]

Sp. nov

Valid

Fang in Fang et al.

Ordovician

 China

A member of Lituitida.

Xiaohenautilus mulleni[26]

Sp. nov

Valid

Jenks et al.

Early Triassic

Dinwoody Formation

 United States
( Nevada)

A member of Nautilida belonging to the group Trigonoceratoidea and the family Grypoceratidae.

Research[edit]

  • New fossil material with shell features that characterize early cephalopods is described from the Cambrian (Terreneuvian) Bonavista Formation (Canada) by Hildenbrand et al. (2021), who interpret this finding as possibly representing the earliest cephalopods known to date, potentially pushing the origin of cephalopods back in time by about 30 million years, to the time before the first occurrence of euarthropods.[63]
  • Two specimens of Syrionautilus libanoticus preserving soft parts, including a specimen showing an imprint of the conch, are described from the Cenomanian of Hadjoula (Lebanon) by Klug et al. (2021).[64]
  • A study on the diversity patterns and spatial structure of belemnite assemblages in Western Tethys Ocean during the Early Jurassic is published by Neige, Weis & Fara (2021).[65]
  • A study on morphological changes of belemnites across the Pliensbachian-Toarcian boundary event, based on data from the Peniche GSSP section (Portugal), is published by Nätscher et al. (2021).[66]
  • A study on chemistry, organization and genesis of circular structures (superficially resembling chromatophores) preserved in coleoid cephalopod specimens from the Jurassic of Germany and the Cretaceous of Lebanon is published by Klug et al. (2021).[67]
  • A well-preserved soft-body imprint of a fossil squid is described from the lower Oligocene of the Krasnodar region (Russia) by Mironenko et al. (2021), who interpret this finding as the first unquestionable representative of Teuthida in the fossil record known to date, and report preservation of fish remains in the stomach contents of this squid, indicative of its piscivorous diet.[68]
  • A specimen of Jeletzkyteuthis coriacea holding a specimen of Parabelopeltis flexuosa in its arms, with the apex very close to its jaws, is described from the Toarcian Posidonia Shale (Germany) by Klug et al. (2021), who interpret this finding as a case of frozen predatory behaviour of two different octobrachian species.[69]
  • Redescription of the gladius of the holotype specimen of Necroteuthis hungarica is published by Košťák et al. (2021), who reinterpret this species as a member of the family Vampyroteuthidae linking Mesozoic loligosepiids with extant Vampyroteuthis.[70]
  • Klompmaker & Landman (2021) describe the oldest drill holes produced by octopodoids in lucinid bivalve specimens from the Campanian of South Dakota, indicating that the drilling habit evolved early in the evolutionary history of Octopodoidea.[71]

Gastropods[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Acirsa ambrosii[72]

Sp. nov

Valid

Brunetti & Cresti

Pliocene

 Italy

A species of Acirsa.

Acirsa (Notacirsa) texana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Acirsa.

Acteon pseudotypica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Acteon.

Acutitomaria[74]

Gen. et sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida. Genus includes new species A. kustatscherae.

Alatostrombus[75]

Gen. et sp. et comb. nov

Valid

Raven

Miocene

 Indonesia
 Malaysia

A member of the family Strombidae. Genus includes new species A. acutalatus, as well as "Strombus" sedanensis Martin (1899).

Alvania absuturalis[76]

Sp. nov

Valid

Chirli & Forli

Pliocene

 Italy

A species of Alvania.

Alvania ameliae[76]

Sp. nov

Valid

Chirli & Forli

Pleistocene

 Italy

A species of Alvania.

Alvania benestarensis[76]

Sp. nov

Valid

Chirli & Forli

Miocene

 Italy

A species of Alvania.

Alvania cerreti[77]

Sp. nov

Valid

Gardella et al.

Pliocene

 Italy

A species of Alvania.

Alvania pluricosticillata[76]

Sp. nov

Disputed

Chirli & Forli

Plio-Pleistocene

 Italy

Originally described as a species of Alvania. Argued to be a junior synonym of the extant taxon Crisilla gagliniae by Amati & Oliverio (2024).[78]

Amaea (Confusiscala) paleocaenica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Ammonicera danieli[79]

Sp. nov

Valid

Chernyshev & Goedert

Late Eocene/early Oligocene

Gries Ranch Formation

 United States
( Washington)

A species of Ammonicera.

Ammonicera rolani[79]

Sp. nov

Valid

Chernyshev & Goedert

Late Eocene/early Oligocene

Gries Ranch Formation

 United States
( Washington)

A species of Ammonicera.

Ampezzogyra angulata[80]

Sp. nov

Valid

Nützel & Hausmann in Hausmann et al.

Late Triassic (Carnian)

San Cassiano Formation

 Italy

A member of the family Stuoraxidae.

Amphidromus ubaldii[81]

Sp. nov

Valid

Dharma

Late Pliocene

 Indonesia

A species of Amphidromus.

Amplitomaria[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" spuria Münster (1841).

Anacithara akisi[82]

Sp. nov

Valid

Kolokotronis

Pliocene

 Cyprus

A species of Anacithara.

Anceps siminescui[83]

Nom. nov

Valid

Harzhauser

Middle-late Miocene

 Moldova

A Trochidae member;
a replacement name for Trochus semistriatus Siminescu & Barbu (1940).

Angulatella[80]

Gen. et sp. nov

Valid

Nützel & Hausmann in Hausmann et al.

Late Triassic (Carnian)

San Cassiano Formation

 Italy

A member of the family Prostyliferidae. The type species is A. bizzarinii.

Antithala[84]

Gen. et comb. et 2 sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Bosnia and Herzegovina
 Hungary
 Poland
 Romania
 Ukraine

A member of the family Costellariidae. The type species is "Mitra" sturi Hoernes & Auinger (1880); genus also includes "Mitra" neugeboreni Hoernes & Auinger (1880), as well as new species A. claviformis and A. filipescui.

Apiocypraea cerretensis[85]

Sp. nov

Valid

Fehse et al.

Pliocene (Zanclean)

 Italy

A member of the family Eocypraeidae.

Aplus abessensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Aplus.

Aplus grandiculus[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Aplus.

Aplus littoralis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Aplus.

Aplus pseudoaquitanensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Aplus.

Aquilofusus? antiqua[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Ascheria canni[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A member of the family Hokkaidoconchidae.

Asiaharpa[88]

Gen. et sp. nov

Valid

Raven

Miocene

 Malaysia

A member of the family Harpidae. Genus includes new species A. sarawakiana.

Attenuata? danica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Possibly a species of Attenuata.

Auingeria[89]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Czech Republic
 Germany
 Hungary

A member of the family Columbellidae. The type species is "Columbella" bellardii Hörnes (1852); genus also includes "Columbella (Anachis)" austriaca Hoernes & Auinger (1880) and "Pyrene (Anachis) bellardii" grussbachensis Csepreghy-Meznerics (1969; raised to the rank of a separate species Auingeria grussbachensis).

Axonella[90]

Gen. et comb. et sp. nov

Valid

Pacaud, Lhomme & Renaud

Eocene (Ypresian)

 France
 Spain

A member of the family Hemisinidae. The type species is "Cerithium" palense Rouault (1850); genus also includes "Potamides" granosus Doncieux (1908), as well as new species A. falsiacum.

Babylonia chattica[91]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Babylonia.

Babylonia edigittenbergeri[92]

Sp. nov

Valid

Raven

Miocene

Miri Formation

 Malaysia

A species of Babylonia.

Babylonia sarawakiana[92]

Sp. nov

Valid

Raven

Miocene

Miri Formation

 Malaysia

A species of Babylonia.

Bandellina compacta[80]

Sp. nov

Valid

Nützel & Hausmann in Hausmann et al.

Late Triassic (Carnian)

San Cassiano Formation

 Italy

A member of the family Cornirostridae.

Barycypraea abbasi[93]

Sp. nov

Valid

Dovesi & Parsons

Miocene (Serravallian)

Nyalindung Formation

 Indonesia

A species of Barycypraea.

Barycypraea alessandrodovesi[93]

Sp. nov

Valid

Dovesi & Parsons

Miocene (Serravallian)

Nyalindung Formation

 Indonesia

A species of Barycypraea.

Bela pediensis[94]

Sp. nov

Valid

Kolokotronis

Pliocene

 Cyprus

A species of Bela.

Bellacolumbella[89]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Czech Republic
 Hungary
 Romania

A member of the family Columbellidae. The type species is "Columbella (Nitidella)" karreri Hoernes & Auinger (1880); genus also includes "Columbella (Nitidella)" embryonalis Boettger (1902) and "Columbella (Nitidella)" katharinae Hoernes & Auinger (1880).

Bellardithala[84]

Nom. et 4 sp. nov

Valid

Harzhauser & Landau

Miocene and Pliocene

Baden Formation
Dej Formation

 Austria
 Czech Republic
 Italy
 Poland
 Romania
 Ukraine
 Hungary?

A member of the family Costellariidae; a replacement name for Micromitra Bellardi (1888). The type species is "Voluta" obsoleta Brocchi (1814); genus also includes new species B. kovaci, B. baluki, B. fedosovi and B. dacica.

Bieleraxis[73]

Gen. et 2 sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species B. discoidea and B. altis.

Bithynia disregularis[95]

Sp. nov

In press

Li et al.

Oligocene

Yehucheng Formation

 China

A species of Bithynia.

Bithynia obliquus[95]

Sp. nov

In press

Li et al.

Oligocene

Yehucheng Formation

 China

A species of Bithynia.

Bithynia parainclinimarga[95]

Sp. nov

In press

Li et al.

Oligocene

Yehucheng Formation

 China

A species of Bithynia.

Bithynia paramonolithic[95]

Sp. nov

In press

Li et al.

Oligocene

Yehucheng Formation

 China

A species of Bithynia.

Bittium prekoeneni[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Bittium.

Bulimulus frenguellii[96]

Sp. nov

Valid

Cabrera et al.

Late Cretaceous

Queguay Formation

 Uruguay

A species of Bulimulus.

Burminella[97]

Gen. et sp. nov

In press

Balashov, Perkovsky & Vasilenko

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A land snail possessing morphological traits intermediate between members of the family Pupinidae and other members of Cyclophoroidea. Genus includes new species B. artiukhini.

Bursa transeuntis[98]

Sp. nov

Valid

Forli & Smriglio

Miocene and Pliocene

 Hungary
 Italy
 Poland

A species of Bursa.

Caecum texanum[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Caecum.

Califrapana[99]

Gen. et comb. nov

Valid

Powell & Houart

Late Oligocene to early Miocene

Vaqueros Formation

 Mexico
 United States
( California)

A member of the family Muricidae. The type species is "Purpura" vaquerosensis Arnold (1907).

Calyptraphorus velusus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Camponaxis bandeli[100]

Sp. nov

Valid

Pieroni, Monari & Todd

Late Triassic (Carnian)

San Cassiano Formation

 Italy

A Tofanellidae member

Campylacrum texanum[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Canariella (Majorata) gerti[101]

Sp. nov

Valid

Groh

Quaternary

 Canary Islands

A species of Canariella.

Cancellotomaria[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" subcancellata d'Orbigny (1850).

Cancilla nanostriatula[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Bulgaria
 Hungary

A species of Cancilla.

Cancilla wagreichi[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

A species of Cancilla.

Capulus gelus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Capulus.

Cassis kalmani[103]

Sp. nov

Valid

Vicián & Kovács

Eocene (Lutetian)

Csernye Formation

 Hungary

A species of Cassis.

Cassis rasae[104]

Sp. nov

Valid

Petuch & Berschauer

Pleistocene

 United States
( Florida)

A species of Cassis.

Cassis viliusi[104]

Sp. nov

Valid

Petuch & Berschauer

Pleistocene

 United States
( Florida)

A species of Cassis.

Cenorhytis[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species C. groesbecki.

Ceratopea? moridunensis[105]

Sp. nov

Valid

Ebbestad & Cope

Ordovician (Floian)

 United Kingdom

A member of the family Raphistomatidae.

Cerithiella acuta[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Cerithiella.

Cerithiscala parvibrazosa[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Cerithiscala tehuacana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Chemnitzia silvai[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Chemnitzia.

Chevallieria texanopsis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Chicomurex parvus[107]

Sp. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Chicomurex.

Chicoreus javanus[107]

Sp. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Chicoreus.

Chicoreus martini[107]

Sp. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Chicoreus.

Chicoreus solidus[107]

Sp. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Chicoreus.

Clava (Semivertagus) scriba[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Clavatula hirmetzli[108]

Sp. nov

Valid

Kovács & Vicián

Middle Miocene

Lajta Limestone Formation

 Hungary

A species of Clavatula.

Clavatula santhai[108]

Sp. nov

Valid

Kovács & Vicián

Middle Miocene

Lajta Limestone Formation

 Hungary

A species of Clavatula.

Clavatula szekelyhidiae[108]

Sp. nov

Valid

Kovács & Vicián

Middle Miocene

Lajta Limestone Formation

 Hungary

A species of Clavatula.

Cochlespira (Tahusyrinx) laevis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Cochlespira.

Conilithes herodus[109]

Sp. nov

Valid

Psarras, Koskeridou & Merle

Miocene (Tortonian)

 Greece

A species of Conilithes.

Conradconfusus nodulinus[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

Conradconfusus texanus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Conus (Kalloconus) asterousiaensis[109]

Sp. nov

Valid

Psarras, Koskeridou & Merle

Miocene (Tortonian)

 Greece
 Italy

A species of Conus.

Conus (Kalloconus) helladicus[109]

Sp. nov

Valid

Psarras, Koskeridou & Merle

Miocene (Tortonian)

 Greece

A species of Conus.

Cordieria biouesensis[111]

Sp. nov

Valid

Pacaud

Paleocene (Thanetian)

 France

A species of Cordieria.

Coronia argilla[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Costoanachis venzoi[89]

Nom. nov

Valid

Harzhauser & Landau

Miocene

 Italy

A species of Costoanachis; a replacement name for Anachis (Costoanachis) subcorrugata Venzo & Pelosio (1963).

Crassaseila[73]

Gen. et 2 sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species C. ripleyana and C. paleocenica.

Crassispira plioibericostricta[112]

Sp. nov

Valid

Brunetti & Forli

Pliocene (Zanclean)

 Spain

A species of Crassispira.

Crassispira pseudoglabra[112]

Sp. nov

Valid

Brunetti & Forli

Pliocene (Piacenzian)

 Italy

A species of Crassispira.

Crenaturricula minuta[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Creonella hebetertia[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Cretatortulosa cretakachinensis[113]

Sp. nov

In press

Yu, Salvador & Jarzembowski

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Cretatortulosa gignens[114][115][116]

Sp. nov

In press

Jochum, Yu & Neubauer

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Pupinidae.

Cryptochorda paleocaenica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Cyathopoma pingyiensis[117]

Sp. nov

In press

Yu et al.

CretaceousPaleogene transition

Pingyi Basin

 China

A species of Cyathopoma.

Cylichna (Cylichnopsis) elliptica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Cylichna.

Cymatiella dulaii[103]

Sp. nov

Valid

Vicián & Kovács

Eocene (Lutetian)

Csernye Formation

 Hungary

A species of Cymatiella.

Cymatiella tzankovi[103]

Nom. nov

Valid

Vicián & Kovács

Paleocene

 Bulgaria

A species of Cymatiella; a replacement name for Eutritonium (Sassia) rutoti Tzankov (1940).

Cymatium danianum[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Cymatium.

Cyphoma guraboensis[118]

Sp. nov

Valid

Groves & Landau

Early Pliocene

Gurabo Formation

 Dominican Republic

A species of Cyphoma.

Cypraeorbis alisonkayae[118]

Sp. nov

Valid

Groves & Landau

Miocene

 Dominican Republic

A species of Cypraeorbis.

Cyprioconcha[87]

Gen. et sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A member of the family Hokkaidoconchidae. The type species is C. robertsoni.

Defensina[89]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

 Bosnia and Herzegovina
 Bulgaria
 Romania

A member of the family Columbellidae. The type species is "Columbella (Mitrella)" bucciniformis Hoernes & Auinger (1880).

Dermomurex wanneri[107]

Nom. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Dermomurex; a replacement name for Murex acuticostatus Wanner & Hahn (1935).

Desbruyeresia kambiaensis[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A species of Desbruyeresia.

Desbruyeresia kinousaensis[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A species of Desbruyeresia.

Desbruyeresia memiensis[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A species of Desbruyeresia.

Discotectonica paracutissima[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Discotectonica.

Doliella parnitens[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Doliella.

Domiporta amoena[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Austria
 Bulgaria
 Romania

A species of Domiporta.

Domiporta pulchra[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

A species of Domiporta.

Domiporta turpis[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria

A species of Domiporta.

Ederazyga[100]

Gen. et sp. et comb. nov

Valid

Pieroni, Monari & Todd

Late Triassic (Carnian to Rhaetian)

Nayband Formation
San Cassiano Formation
Zu Limestone

 Iran
 Italy
 Slovenia

A possible Zygopleuridae member.
The type species is E. fanchini;
genus also includes "Cerithium"? lateplicatum Klipstein (1843).

Ellobium mizutanii[119]

Sp. nov

Valid

Kawase & Ichihara

Early Miocene

Akeyo Formation

 Japan

A species of Ellobium.

Entemnotrochus kathiawarensis[120]

Sp. nov

Valid

Bose, Das & Mondal

Miocene (Burdigalian)

 India

A species of Entemnotrochus.

Eoacteon paracutissima[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Eoancilla lapicidina[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Eocithara (?) laurencei[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Eopleurotoma atacica[111]

Nom. nov

Valid

Pacaud

Eocene (Ypresian)

 France

A species of Eopleurotoma; a replacement name for Pleurotoma (Eopleurotoma) romani Doncieux (1908).

Eotrivia pacaudi[121]

Sp. nov

Valid

Fehse

Eocene

 Italy

Eoturris fallsensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Eoworthenia frydai[74]

Nom. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida; a replacement name for Worthenia rarissima Barrande.

Episcomitra antibellardii[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

A species of Episcomitra.

Episcomitra leopoldiana[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Czech Republic
 Poland

A species of Episcomitra.

Episcomitra missile[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Grund Formation

 Austria
 Hungary

A species of Episcomitra.

Episcomitra neubaueri[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

A species of Episcomitra.

Episcomitra pontonxensis[91]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Episcomitra.

Episcomitra pseudoincognita[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania
 Ukraine

A species of Episcomitra.

Episcomitra vermeuleni[122]

Sp. nov

Valid

Pacaud

Eocene (Priabonian)

 Ukraine

A species of Episcomitra.

Eulimella ariejansseni[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Eulimella.

Euthema myanmarica[123]

Sp. nov

Valid

Balashov

Cretaceous

Hkamti amber

 Myanmar

A species of Euthema.

Euthria mellianensis[86]

Sp. nov

Valid

Lozouet

Miocene

 France

A species of Euthria.

Euthria tarusatensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Euthria.

Euthriofusus argillensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

Falsifusus problematicus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Fedosovia[84]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

 Czech Republic
 Italy
 Romania

A member of the family Costellariidae. The type species is "Mitra (Callithea)" fuchsi Hoernes & Auinger (1880).

Fraudiziba[102]

Gen. et nom. et sp. nov

Valid

Harzhauser & Landau

Miocene and Pliocene

Karaman Basin

 Austria
 Bosnia and Herzegovina
 Czech Republic
 Hungary
 Italy
 Poland
 Romania
 Slovakia
 Turkey
 Ukraine

A member of the family Mitridae. The type species is F. paratethyca (a replacement name for Mitra austriaca Csepreghy-Meznerics, 1950); genus also includes new species F. ottomanica.

Fulgurofusus spinosus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Fulgurofusus.

Fusulculus texanus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Fusulculus.

Galba prima[124]

Sp. nov

Valid

Yu, Neubauer & Jochum

Cretaceous

Burmese amber

 Myanmar

A species of Galba.

Gemmula bearrizensis[111]

Nom. nov

Valid

Pacaud

Eocene (Priabonian)

 France

A species of Gemmula; a replacement name for Drillia pellati Boussac (1911).

Gemmula osca[111]

Sp. nov

Valid

Pacaud

Eocene (Bartonian)

 Spain

A species of Gemmula.

Gentilifulgur[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species G. praemortoni.

Gibbula lovellreevei[83]

Nom. nov

Valid

Harzhauser

Pliocene

 United Kingdom

A Trochidae member;
a replacement name for Trochus (Gibbula) reevei Harmer (1923).

Gibbula pizzolatoi[125]

Sp. nov

Valid

Spadini

Pliocene

 Italy

A species of Gibbula.

Gibbula steiningeri[83]

Nom. nov

Valid

Harzhauser

Early Miocene

 Austria

A Trochidae member;
a replacement name for Trochus amedei bicincta Schaffer (1912).

Gibbula tavanii[83]

Nom. nov

Valid

Harzhauser

Miocene

 Libya

A Trochidae member;
a replacement name for Gibbula minima Tavani (1939).

Gibbuliculus[83]

Gen. et comb. et nom. nov

Valid

Harzhauser

Oligocene to Pleistocene

 Europe

A Trochidae member;
The type species is "Gibbula" brebioni Landau, Van Dingenen & Ceulemans (2017).
Genus includes G. saccoi (a replacement name for Gibbula protumida Sacco, 1896).

"Gibbulinella" sandbergeri[126]

Nom. nov

Valid

Harzhauser & Neubauer

Eocene

 Italy

A member of the family Streptaxidae; a replacement name for Pupa simplex Sandberger (1870).

Haedropleura dellabellai[72]

Sp. nov

Valid

Brunetti & Cresti

Pliocene

 Italy

A species of Haedropleura.

Hammatospira cancellata[127]

Sp. nov

Valid

Mazaev

Early Carboniferous

 Kazakhstan

Hespererato canae[118]

Sp. nov

Valid

Groves & Landau

Late Miocene and early Pliocene

Cercado Formation

 Dominican Republic

A species of Hespererato.

Hespererato praeclusa[118]

Sp. nov

Valid

Groves & Landau

Early–middle Miocene

Baitoa Formation

 Dominican Republic

A species of Hespererato.

Hexaplex nascium[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Hexaplex.

Hirsuticyclus canaliculatus[128]

Sp. nov

In press

Yu

Cretaceous

Burmese amber

 Myanmar

A member of the family Cyclophoridae.

Hochheimia[126]

Nom. nov

Valid

Harzhauser & Neubauer

Oligocene

 Germany

A member of the family Hygromiidae; a replacement name for Palaeotrichia Nordsieck (2014).

Hokkaidoconcha morisseaui[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A member of the family Hokkaidoconchidae.

Holospira? fallsensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Possibly a species of Holospira.

?Hydrocena praecursor[129]

Sp. nov

In press

Yu & Neubauer

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Hydrocenidae.

Jponsia lacetania[130]

Sp. nov

Valid

Pacaud & Goret

Eocene (Ypresian)

 Spain

A member of the family Pachychilidae.

Kapalmerella gelus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Keepingia elongata[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

Laevispira[75]

Gen. et comb. et sp. nov

Valid

Raven

Miocene and Pliocene

 Brunei
 India
 Indonesia

A member of the family Strombidae. Genus includes "Dolomena" bruneiensis Harzhauser, Raven & Landau (2018), "Strombus" glaber Martin (1879), "Strombus" karikalensis Cossmann (1903), "Strombus" triangulatus Martin (1879) and "Strombus" varinginensis Martin (1899), as well as new species L. seriaensis.

Laevitrivia[131]

Gen. et comb. et 5 sp. nov

Valid

Fehse & Vicián

Miocene and Pliocene

 France
 Italy

A member of the family Triviidae. The type species is "Cypraea" pisolina Lamarck, (1810); genus also includes new species L. annae, L. bertaccinii, L. pliopisolina, L. pseudocalva and L. ventricostata.

Latirus? asperpellis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Possibly a species of Latirus.

Laubella subsulcata[74]

Sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida.

Lineacingulum[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" texturata Münster (1841).

Lineaetomaria[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" decorata Münster (1841).

Liostomia wilvanderstoelae[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Liostomia.

Littorina elongata[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Littorina.

Littorina (Prosthenodon) incertus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Littorina.

Lupira danianum[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Lutema gracilis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Lyria elongata[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Lyria.

Lyria tehuacana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Lyria.

Malea hyaducki[104]

Sp. nov

Valid

Petuch & Berschauer

Pliocene

 United States
( Florida)

A species of Malea.

Martaia[89]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

 Austria
 Bulgaria
 Czech Republic
 Hungary
 Romania
 Ukraine

A member of the family Columbellidae. The type species is "Columbella (Anachis)" dujardini Hoernes & Auinger (1880); genus also includes "Columbella (Anachis)" zitteli Hoernes & Auinger (1880).

Mathilda (Echinimathilda) duogenta[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Mathilda.

Melongena pontonxensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Melongena.

Metula bulbiformis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Metula.

Metula nana[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Metula.

Metula pseudomajor[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Metula.

Metula rivulisensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Metula.

Mimospira llangynogensis[105]

Sp. nov

Valid

Ebbestad & Cope

Ordovician (Floian)

 United Kingdom

A member of the family Clisospiridae.

Mitrella dacica[89]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Hungary
 Romania
 Slovakia

A species of Mitrella.

Mitrella demaintenonae[89]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation
Dej Formation

 Austria
 Romania

A species of Mitrella.

Mitrella elongatissima[89]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Bosnia and Herzegovina

A species of Mitrella.

Mitrella fortis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Mitrella.

Mitrella tarbelliana[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Mitrella.

Mitrella viennensis[89]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Bulgaria
 Czech Republic
 Kazakhstan
 Poland
 Romania

A species of Mitrella.

Moniquia astibiai[130]

Sp. nov

Valid

Pacaud & Goret

Eocene (Priabonian)

 Spain

A member of the family Pachychilidae.

Monoplex s.l. szakonyii[103]

Sp. nov

Valid

Vicián & Kovács

Eocene (Lutetian)

Csernye Formation

 Hungary

A member of the family Cymatiidae.

Mulderia mulderi[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A member of the family Pyramidellidae.

Multitrochia[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species M. tehuacana.

Nannamoria alquezae[132]

Sp. nov

Valid

Hawke

Miocene

 Australia

A species of Nannamoria.

Nannamoria cadella[132]

Sp. nov

Valid

Hawke

Miocene

Cadel Formation

 Australia

A species of Nannamoria.

Nannamoria costatum[132]

Sp. nov

Valid

Hawke

Pliocene

Cameron Inlet Formation

 Australia

A species of Nannamoria.

Nannamoria flindersi[132]

Sp. nov

Valid

Hawke

Pliocene

Cameron Inlet Formation

 Australia

A species of Nannamoria.

Nannamoria gnotuki[132]

Sp. nov

Valid

Hawke

 Australia

A species of Nannamoria.

Nannamoria hiscocki[132]

Sp. nov

Valid

Hawke

Pliocene

Cameron Inlet Formation

 Australia

A species of Nannamoria.

Nannamoria malonei[132]

Sp. nov

Valid

Hawke

 Australia

A species of Nannamoria.

Nannamoria persimilis[132]

Sp. nov

Valid

Hawke

Miocene

 Australia

A species of Nannamoria.

Naria praehelvola[133]

Sp. nov

Valid

Fehse & Vicián

Miocene

 Indonesia

A species of Naria.

Natica (Carinacca) kincaidensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Natica.

Nebularia soliphila[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria

A species of Nebularia.

Nemaspira[134]

Gen. et comb. et 2 sp. nov

Valid

Mazaev

Early Carboniferous to Late Triassic

 Belgium
 Russia
 Thailand
 United Kingdom
 United States
 China?

A member of the family Gosseletinidae. Genus includes several species formerly assigned to the genera Gosseletina and Globodoma, as well as new species N. shamaevi and N. skuini.

Nitidiclavus senesi[135]

Sp. nov

Valid

Biskupič

Miocene

 Slovakia

A member of the family Drilliidae.

Niveria callosa[136]

Sp. nov

Valid

Fehse

Late Oligocene

 France

A species of Niveria.

Niveria novosperara[136]

Sp. nov

Valid

Fehse

Late Oligocene

 France

A species of Niveria.

Nodocingulum[74]

Gen. et comb. et 2 sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida. The type species is "Pleurotomaria" coronata Münster (1841); genus also includes new species N. ernstkittli and possibly also N? turris.

Nodolargena[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A member of the new family Nodolargenidae. Genus includes new species N. frosti.

Nodovoluta[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species N. coronata.

Odostomia malagensis[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Odostomia.

Oligonalia[86]

Gen. et comb. nov

Valid

Lozouet

Oligocene

 France

A new genus for "Siphonalia" riparia Lozouet (1999).

Olinderia[91]

Gen. et comb. et 3 sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Charitodoronidae. The type species is "Charitodoron" tauzini Lozouet (1991); genus also includes new species O. lestrillensis, O. poustagnacensis and O. verdunensis.

Olivella tehuacanensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Olivella.

Ondina pinguis[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Ondina.

Onustus undosus[75]

Sp. nov

Valid

Raven

Miocene

 Brunei

A species of Onustus.

Opalia tehuacana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Opalia.

Orbitestella kieli[79]

Sp. nov

Valid

Chernyshev & Goedert

Late Eocene/early Oligocene

Gries Ranch Formation

 United States
( Washington)

A member of the family Orbitestellidae.

Orthosurcula ethani[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

Oxyacrum gracilis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Pagonia[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species P. spiralis.

Palaeorhaphis? laevis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Paleofusus[73]

Gen. et 3 sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species P. bellus, P. parvicrassus and P. elegantissima.

Papuliscala? acus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation